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© reprodução total
Basso, Thiago Olitta.
Melhoramento da fermentação alcóolica em Saccharomyces
cerevisiae por engenharia evolutiva / Thiago Olitta Basso. -- São
Paulo, 2011.
Orientador: Andreas Karoly Gombert.
Tese (Doutorado) – Universidade de São Paulo. Instituto de Ciências Biomédicas. Programa de Pós-Graduação Interunidades em
Biotecnologia USP/IPT/Instituto Butantan. Área de concentração: Biotecnologia. Linha de pesquisa: Fisiologia e engenharia metabólica de leveduras.
Versão do título para o inglês: Improvement of alcoholic fermentation
in Saccharomyces cerevisiae by evolutionary engineering.
Descritores: 1. Sacarose 2. Saccharomyces cerevisiae 3. Invertase
4. Engenharia evolutiva 5. Etanol 6. AGT1 I. Gombert, Andreas
Karoly II. Universidade de São Paulo. Instituto de Ciências Biomédicas. Programa de Pós-Graduação Interunidades em Biotecnologia
USP/IPT/Instituto Butantan III. Título.
Universidade de São Paulo, Instituto Butantan, Instituto de Pesquisas Tecnológicas
______________________________________________________________________________________________________________
Candidato(a): Thiago Olitta Basso.
Título da Tese: Melhoramento da fermentação alcóolica em Saccharomyces
cerevisiae por engenharia evolutiva.
Orientador(a): Andreas Karoly Gombert.
A Comissão Julgadora dos trabalhos de Defesa da Tese de Doutorado, em sessão pública realizada a .../.../..., considerou
( ) Aprovado(a) ( ) Reprovado(a)
Examinador(a): Assinatura: ... Nome: ... Instituição: ...
Examinador(a): Assinatura: ... Nome: ... Instituição: ...
Examinador(a): Assinatura: ... Nome: ... Instituição: ...
Examinador(a): Assinatura: ... Nome: ... Instituição: ...
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]$++ITTo^O!?*B3S!<>9MJQF5!∆N∆5GEU!]$++ITTp^O!?*B3S!@"@$UI+%(Un!]$+$ITqq^!<!?*B3S!@"/#nI
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"5P<M5! tRq! IIII! (?=E5! F9=! p! G<D<=! 69:! <dC8<==L9! F?:?DJ?F5! D5! M?D75G<:! ?*B3S! <:! 8<M5KL9! l!
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U $."-'/B12'$."-'/B12'$."-'/B12'$."-'/B12' RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR SUSUSUSU
S SS
S -,A$*2'!)$)($'&-V0$3%-,A$*2'!)$)($'&-V0$3%-,A$*2'!)$)($'&-V0$3%RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR-,A$*2'!)$)($'&-V0$3%RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR StRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRStStSt
SRU SRUSRU
SRU ,DG<D758?5!+<E5PHM?65!F<!,DG<D758?5!+<E5PHM?65!F<!,DG<D758?5!+<E5PHM?65!F<!,DG<D758?5!+<E5PHM?65!F<!*R!6<8<>?=?5<*R!6<8<>?=?5<*R!6<8<>?=?5<*R!6<8<>?=?5<!C585!9!:<M7985:<DE9!F5!C89FJKL9!F<!<E5D9M!!C585!9!:<M7985:<DE9!F5!C89FJKL9!F<!<E5D9M!!C585!9!:<M7985:<DE9!F5!C89FJKL9!F<!<E5D9M!!C585!9!:<M7985:<DE9!F5!C89FJKL9!F<!<E5D9M!
69:PJ=EQ><M 69:PJ=EQ><M69:PJ=EQ><M
69:PJ=EQ><MRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR StRRRRRRRRRRRRRRRRRRRRRRStStSt
SRS SRSSRS
SRS %!,DG<D758?5!,>9MJE?>5!69:9!Y<885:<DE5!69:CM<:<DE58!l!,DG<D758?5!+<E5PHM?65%!,DG<D758?5!,>9MJE?>5!69:9!Y<885:<DE5!69:CM<:<DE58!l!,DG<D758?5!+<E5PHM?65%!,DG<D758?5!,>9MJE?>5!69:9!Y<885:<DE5!69:CM<:<DE58!l!,DG<D758?5!+<E5PHM?65%!,DG<D758?5!,>9MJE?>5!69:9!Y<885:<DE5!69:CM<:<DE58!l!,DG<D758?5!+<E5PHM?65 SsSsSsSs
SRn SRnSRn
SRn +<E5P9M?=:9!F<!=56589=<!<:!+<E5P9M?=:9!F<!=56589=<!<:!+<E5P9M?=:9!F<!=56589=<!<:!+<E5P9M?=:9!F<!=56589=<!<:!*R!6<8<>?=?5<*R!6<8<>?=?5<*R!6<8<>?=?5<*R!6<8<>?=?5<RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR nTRRRRRRRRRRRRRRRRRRRRRRRRRRRRnTnTnT
SRnRU SRnRU SRnRU
SRnRU A?5!<dE856<MJM58!<!>?5!?DE856<MJM58A?5!<dE856<MJM58!<!>?5!?DE856<MJM58A?5!<dE856<MJM58!<!>?5!?DE856<MJM58A?5!<dE856<MJM58!<!>?5!?DE856<MJM58RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR nTRRRnTnTnT
SRnRS SRnRS SRnRS
SRnRS "85D=C98E<!5E?>9!F<!=56589=<!5E85>`=!F5!:<:P85D5!CM5=:ZE?65!F<!M<><FJ85="85D=C98E<!5E?>9!F<!=56589=<!5E85>`=!F5!:<:P85D5!CM5=:ZE?65!F<!M<><FJ85="85D=C98E<!5E?>9!F<!=56589=<!5E85>`=!F5!:<:P85D5!CM5=:ZE?65!F<!M<><FJ85="85D=C98E<!5E?>9!F<!=56589=<!5E85>`=!F5!:<:P85D5!CM5=:ZE?65!F<!M<><FJ85=RRRRRRRRRRRRRRRRRRRR
!!!! RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR nUnUnUnU
SRnRn SRnRn SRnRn
SRnRn -<GJM5KL9!GaD?65!F9=!M96?!+%(-<GJM5KL9!GaD?65!F9=!M96?!+%(-<GJM5KL9!GaD?65!F9=!M96?!+%(-<GJM5KL9!GaD?65!F9=!M96?!+%(RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR ntRRRRRRntntnt
SRnRr SRnRr SRnRr
SRnRr -<GJM5KL9!GaD?65!F9=!M96?!*B3-<GJM5KL9!GaD?65!F9=!M96?!*B3-<GJM5KL9!GaD?65!F9=!M96?!*B3-<GJM5KL9!GaD?65!F9=!M96?!*B3RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR nsRRRRRRRnsnsns
SRr SRrSRr
SRr 0?=?9M9G?5! [J5DE?E5E?>5! F<!0?=?9M9G?5! [J5DE?E5E?>5! F<!0?=?9M9G?5! [J5DE?E5E?>5! F<!0?=?9M9G?5! [J5DE?E5E?>5! F<!*R! 6<8<>?=?5<*R! 6<8<>?=?5<*R! 6<8<>?=?5<*R! 6<8<>?=?5<! FJ85DE<! 9! 68<=6?:<DE9! <:! =56589=<! 9J! <:!! FJ85DE<! 9! 68<=6?:<DE9! <:! =56589=<! 9J! <:!! FJ85DE<! 9! 68<=6?:<DE9! <:! =56589=<! 9J! <:!! FJ85DE<! 9! 68<=6?:<DE9! <:! =56589=<! 9J! <:!
:?=EJ85=!F<!GM?69=<!<!Y8JE9=< :?=EJ85=!F<!GM?69=<!<!Y8JE9=<:?=EJ85=!F<!GM?69=<!<!Y8JE9=<
:?=EJ85=!F<!GM?69=<!<!Y8JE9=<RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR nononono
n nn
n ')i,"$A'*')i,"$A'*')i,"$A'*')i,"$A'*RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR rSrSrSrS
r rr
r +%",-$%$*!,!+€"'/'*+%",-$%$*!,!+€"'/'*+%",-$%$*!,!+€"'/'* RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR+%",-$%$*!,!+€"'/'*RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR rrRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRrrrrrr
rRU rRUrRU
rRU (?D75G<D=!F<!M<><FJ85!<!C8<=<8>5KL9(?D75G<D=!F<!M<><FJ85!<!C8<=<8>5KL9(?D75G<D=!F<!M<><FJ85!<!C8<=<8>5KL9 RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR(?D75G<D=!F<!M<><FJ85!<!C8<=<8>5KL9RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR rrrrrrrr
rRS rRSrRS
rRS +<?9=!F<!6JME?>9+<?9=!F<!6JME?>9+<?9=!F<!6JME?>9+<?9=!F<!6JME?>9RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR rtrtrtrt
rRn rRnrRn
rRn 39DF?Kc<=!F<!6JME?>939DF?Kc<=!F<!6JME?>939DF?Kc<=!F<!6JME?>9 RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR39DF?Kc<=!F<!6JME?>9RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR rqrqrqrq
rRnRU rRnRU rRnRU
rRnRU 3JME?>93JME?>93JME?>93JME?>9!<:!Y85=69!5G?E5F9!<:!Y85=69!5G?E5F9!<:!Y85=69!5G?E5F9!<:!Y85=69!5G?E5F9RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR rqrqrqrq
rRnRS rRnRS rRnRS
rRnRS 3JME?>9=!<:![J?:?9=E5E93JME?>9=!<:![J?:?9=E5E93JME?>9=!<:![J?:?9=E5E93JME?>9=!<:![J?:?9=E5E9RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR rsRRRRRRRRRRRRRRRRrsrsrs
rRnRn rRnRn rRnRn
rRnRn 3JME?>9=!<:!P5E<M5F5=!8<C<E?F5=!]*)-^3JME?>9=!<:!P5E<M5F5=!8<C<E?F5=!]*)-^3JME?>9=!<:!P5E<M5F5=!8<C<E?F5=!]*)-^3JME?>9=!<:!P5E<M5F5=!8<C<E?F5=!]*)-^RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR rprprprp
rRr rRrrRr
rRr /<E<8:?D5KL9!F5!Y85KL9!:9M58!F<!'/<E<8:?D5KL9!F5!Y85KL9!:9M58!F<!'/<E<8:?D5KL9!F5!Y85KL9!:9M58!F<!'/<E<8:?D5KL9!F5!Y85KL9!:9M58!F<!'SSSS!<!3'!<!3'!<!3'S!<!3'SSS!D9!GZ=!F<!=5QF5!F9!P?988<5E98!D9!GZ=!F<!=5QF5!F9!P?988<5E98!D9!GZ=!F<!=5QF5!F9!P?988<5E98RRRRRRRRRRRRRR!D9!GZ=!F<!=5QF5!F9!P?988<5E98RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tTtTtTtT
rRt rRtrRt
rRt mJ5DE?Y?65KL9!F5!69D6<DE85KL9!F<!:<E5PHM?E9=!<dE856<MJM58<=!<!?DE856<MJMmJ5DE?Y?65KL9!F5!69D6<DE85KL9!F<!:<E5PHM?E9=!<dE856<MJM58<=!<!?DE856<MJMmJ5DE?Y?65KL9!F5!69D6<DE85KL9!F<!:<E5PHM?E9=!<dE856<MJM58<=!<!?DE856<MJMmJ5DE?Y?65KL9!F5!69D6<DE85KL9!F<!:<E5PHM?E9=!<dE856<MJM58<=!<!?DE856<MJM58<=58<=58<=58<= RRRRRRRRRRRRRRRRRR tTRRRRRRtTtTtT
rRq rRqrRq
rRq mJ5DE?Y?65KL9!F5!69D6<DE85KL9!6<MJM58mJ5DE?Y?65KL9!F5!69D6<DE85KL9!6<MJM58mJ5DE?Y?65KL9!F5!69D6<DE85KL9!6<MJM58mJ5DE?Y?65KL9!F5!69D6<DE85KL9!6<MJM58 RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tUtUtUtU
rRs rRsrRs
rRs $D=C<KL9!F5!CJ8<_5!F5!6JMEJ85$D=C<KL9!F5!CJ8<_5!F5!6JMEJ85$D=C<KL9!F5!CJ8<_5!F5!6JMEJ85 RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR$D=C<KL9!F5!CJ8<_5!F5!6JMEJ85RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tStStStS
rRo rRorRo
rRo +<F?F5!F5!65C56?F5F<!Y<8:<DE5E?>5+<F?F5!F5!65C56?F5F<!Y<8:<DE5E?>5+<F?F5!F5!65C56?F5F<!Y<8:<DE5E?>5 RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR+<F?F5!F5!65C56?F5F<!Y<8:<DE5E?>5RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tStStStS
rRp rRprRp
rRp /<E<8:?D5KL9!F5!5E?>?F5F<!F<!?D><8E5=<!<dE856<MJM58!<!E9E5M/<E<8:?D5KL9!F5!5E?>?F5F<!F<!?D><8E5=<!<dE856<MJM58!<!E9E5M/<E<8:?D5KL9!F5!5E?>?F5F<!F<!?D><8E5=<!<dE856<MJM58!<!E9E5M/<E<8:?D5KL9!F5!5E?>?F5F<!F<!?D><8E5=<!<dE856<MJM58!<!E9E5M RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tStStStS
rRUT rRUTrRUT
rRUT /<E<8:?D5KL9!F5!5E?>?F5F<!F9!E85D=C98E<!F<!=56589=</<E<8:?D5KL9!F5!5E?>?F5F<!F9!E85D=C98E<!F<!=56589=</<E<8:?D5KL9!F5!5E?>?F5F<!F9!E85D=C98E<!F<!=56589=</<E<8:?D5KL9!F5!5E?>?F5F<!F9!E85D=C98E<!F<!=56589=<RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tntntntn
rRUU rRUUrRUU
rRUU +5D?CJM5Kc<=!G<D`E?65=+5D?CJM5Kc<=!G<D`E?65=+5D?CJM5Kc<=!G<D`E?65= RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR+5D?CJM5Kc<=!G<D`E?65=RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tttttttt
rRUURU rRUURU rRUURU
rRUURS rRUURS rRUURS
rRUURS -<5KL9!<:!65F<?5!F5!C9M?:<85=<!]@3-^-<5KL9!<:!65F<?5!F5!C9M?:<85=<!]@3-^-<5KL9!<:!65F<?5!F5!C9M?:<85=<!]@3-^-<5KL9!<:!65F<?5!F5!C9M?:<85=<!]@3-^RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tttttttt
rRUURn rRUURn rRUURn
rRUURn "85D=Y98:5KL9!F<!M<><FJ85=!C<M9!:`E9F9!F9!56<E5E9!F<!MQE?9"85D=Y98:5KL9!F<!M<><FJ85=!C<M9!:`E9F9!F9!56<E5E9!F<!MQE?9"85D=Y98:5KL9!F<!M<><FJ85=!C<M9!:`E9F9!F9!56<E5E9!F<!MQE?9"85D=Y98:5KL9!F<!M<><FJ85=!C<M9!:`E9F9!F9!56<E5E9!F<!MQE?9RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tttttttt
rRUURr rRUURr rRUURr
rRUURr -<69D=E?EJ?KL9!F9!G<D<!B-%n-<69D=E?EJ?KL9!F9!G<D<!B-%n-<69D=E?EJ?KL9!F9!G<D<!B-%n-<69D=E?EJ?KL9!F9!G<D<!B-%nRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tqtqtqtq
rRUURt rRUURt rRUURt
rRUURt /<M<KL9!F9!G<D<!%&"U/<M<KL9!F9!G<D<!%&"U/<M<KL9!F9!G<D<!%&"U/<M<KL9!F9!G<D<!%&"URRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR tstststs
rRUURq rRUURq rRUURq
rRUURq /<M<KL9!F9!G<D<!+%(Un/<M<KL9!F9!G<D<!+%(Un/<M<KL9!F9!G<D<!+%(Un/<M<KL9!F9!G<D<!+%(UnRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR qURRRRRRRRRRRRRRRRqUqUqU
rRUURs rRUURs rRUURs
rRUURs *JP=E?EJ?KL9!F9!C89:9E98!F9!G<D<!+%(Un*JP=E?EJ?KL9!F9!C89:9E98!F9!G<D<!+%(Un*JP=E?EJ?KL9!F9!C89:9E98!F9!G<D<!+%(Un*JP=E?EJ?KL9!F9!C89:9E98!F9!G<D<!+%(UnRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR qSqSqSqS
rRUURo rRUURo rRUURo
rRUURo *JC<8*JC<8*JC<8*JC<8IIII<dC8<==L9!F9!G<D<!%&"U<dC8<==L9!F9!G<D<!%&"U<dC8<==L9!F9!G<D<!%&"U<dC8<==L9!F9!G<D<!%&"URRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR qnqnqnqn
rRUS rRUSrRUS
rRUS %DZM?=<!F5!<dC8<==L9!GaD?65!<:!<=65M5!G<D•:?65!]E85D=68?CE9:5^%DZM?=<!F5!<dC8<==L9!GaD?65!<:!<=65M5!G<D•:?65!]E85D=68?CE9:5^%DZM?=<!F5!<dC8<==L9!GaD?65!<:!<=65M5!G<D•:?65!]E85D=68?CE9:5^%DZM?=<!F5!<dC8<==L9!GaD?65!<:!<=65M5!G<D•:?65!]E85D=68?CE9:5^ RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR qrRRRRRRRRRRRRRRRRRRRRRRRRRqrqrqr
rRUn rRUnrRUn
rRUn *<[J<D6?5:<DE9!F9!G<D<!*<[J<D6?5:<DE9!F9!G<D<!*<[J<D6?5:<DE9!F9!G<D<!*<[J<D6?5:<DE9!F9!G<D<!%&"U%&"U%&"U%&"URRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR qtRRRRRRRRRRRRRqtqtqt
rRUr rRUrrRUr
rRUr 3ZM6JM9!F9=!C58{:<E89=!Y?=?9MHG?69=3ZM6JM9!F9=!C58{:<E89=!Y?=?9MHG?69=3ZM6JM9!F9=!C58{:<E89=!Y?=?9MHG?69= RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR3ZM6JM9!F9=!C58{:<E89=!Y?=?9MHG?69=RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR qtRRRRRRRqtqtqt
t tt
t -,*B("%/'*!,!/$*3B**2'-,*B("%/'*!,!/$*3B**2'-,*B("%/'*!,!/$*3B**2' RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR-,*B("%/'*!,!/$*3B**2'RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR qsqsqsqs
tRU tRUtRU
tRU ,Y<?E9=!F5!M?:,Y<?E9=!F5!M?:,Y<?E9=!F5!M?:,Y<?E9=!F5!M?:?E5KL9!C98!J856?M5!<:!6JME?>9=!69DEQDJ9=!F<!?E5KL9!C98!J856?M5!<:!6JME?>9=!69DEQDJ9=!F<!?E5KL9!C98!J856?M5!<:!6JME?>9=!69DEQDJ9=!F<!?E5KL9!C98!J856?M5!<:!6JME?>9=!69DEQDJ9=!F<!*R!6<8<>?=?5<*R!6<8<>?=?5<*R!6<8<>?=?5<*R!6<8<>?=?5<RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR qpRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRqpqpqp
tRS tRStRS
tRS ,=EJF9!F5!8<M965M?_5KL9!F9!:<E5P9M?=:9!F5!=56589=<!<:!6JME?>9=!69DEQDJ9=,=EJF9!F5!8<M965M?_5KL9!F9!:<E5P9M?=:9!F5!=56589=<!<:!6JME?>9=!69DEQDJ9=,=EJF9!F5!8<M965M?_5KL9!F9!:<E5P9M?=:9!F5!=56589=<!<:!6JME?>9=!69DEQDJ9=,=EJF9!F5!8<M965M?_5KL9!F9!:<E5P9M?=:9!F5!=56589=<!<:!6JME?>9=!69DEQDJ9= RRRRRRRRRRRRRRRRRRRRRRRRRRRRRR srRRRRRRRRRRsrsrsr
tRSRU tRSRU tRSRU
tRSRU +9F<M9!E<H8?69!F9!:<E5P9M?=:9!F<!=56589=<!<:!5D5<89P?9=<+9F<M9!E<H8?69!F9!:<E5P9M?=:9!F<!=56589=<!<:!5D5<89P?9=<+9F<M9!E<H8?69!F9!:<E5P9M?=:9!F<!=56589=<!<:!5D5<89P?9=<+9F<M9!E<H8?69!F9!:<E5P9M?=:9!F<!=56589=<!<:!5D5<89P?9=<RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR srsrsrsr
tRSRS tRSRS tRSRS
tRSRS 0?=?9M9G?5!F5=!M?D75G<D=!?*B3S!J850?=?9M9G?5!F5=!M?D75G<D=!?*B3S!J850?=?9M9G?5!F5=!M?D75G<D=!?*B3S!J850?=?9M9G?5!F5=!M?D75G<D=!?*B3S!J85IIII!<!*B3S!J85!<!*B3S!J85!<!*B3S!J85!<!*B3S!J85IIII!<:![J?:?9=E5E9=!M?:?E5F9=!C98!!<:![J?:?9=E5E9=!M?:?E5F9=!C98!!<:![J?:?9=E5E9=!M?:?E5F9=!C98!!<:![J?:?9=E5E9=!M?:?E5F9=!C98!
=56589=<!9J!GM?69=<!<:!5<89 =56589=<!9J!GM?69=<!<:!5<89 =56589=<!9J!GM?69=<!<:!5<89
=56589=<!9J!GM?69=<!<:!5<89IIII!!!!9J!5D5<89P?9=<9J!5D5<89P?9=<9J!5D5<89P?9=<9J!5D5<89P?9=<RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR ssRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRssssss
tRn tRntRn
tRn 39:C585KL9!F5=!M?D75G<D=!39:C585KL9!F5=!M?D75G<D=!39:C585KL9!F5=!M?D75G<D=!39:C585KL9!F5=!M?D75G<D=!*B3S*B3S*B3S*B3S!<!?!<!?!<!?!<!?*B3S*B3S*B3S*B3S!<:!6JME?>9=!F<=69DEQDJ9=!<:!6JME?>9=!F<=69DEQDJ9=!<:!6JME?>9=!F<=69DEQDJ9=!<:!6JME?>9=!F<=69DEQDJ9=RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR oqoqoqoq
tRnRU tRnRU tRnRU
tRnRU 3JME?>9=!F<=69DEQDJ9=!<:!Y85=69!5G3JME?>9=!F<=69DEQDJ9=!<:!Y85=69!5G3JME?>9=!F<=69DEQDJ9=!<:!Y85=69!5G3JME?>9=!F<=69DEQDJ9=!<:!Y85=69!5G?E5F9?E5F9?E5F9?E5F9RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR oqoqoqoq
tRnRS tRnRS tRnRS
tRnRS 3JME?>9=!F<=69DEQDJ9=!<:!5D5<89P?9=<!CM<D53JME?>9=!F<=69DEQDJ9=!<:!5D5<89P?9=<!CM<D53JME?>9=!F<=69DEQDJ9=!<:!5D5<89P?9=<!CM<D53JME?>9=!F<=69DEQDJ9=!<:!5D5<89P?9=<!CM<D5RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR opopopop
tRr tRrtRr
tRr ,DG<D758?5! <>9MJE?>5! C585! 9! :<M7985:<DE9! F9! E85D=C98E<! F<! =5658,DG<D758?5! <>9MJE?>5! C585! 9! :<M7985:<DE9! F9! E85D=C98E<! F<! =5658,DG<D758?5! <>9MJE?>5! C585! 9! :<M7985:<DE9! F9! E85D=C98E<! F<! =5658,DG<D758?5! <>9MJE?>5! C585! 9! :<M7985:<DE9! F9! E85D=C98E<! F<! =56589=<! <:!9=<! <:!9=<! <:!9=<! <:! *R!*R!*R!*R!
6<8<>?=?5< 6<8<>?=?5<6<8<>?=?5<
6<8<>?=?5<RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR ptRRRRRRRRRRRRRRRRRRRRRRRRRptptpt
tRrRU tRrRU tRrRU
tRrRU 0?=?9M9G?5!F<!*R!6<8<>?=?5<!*B3S!<!?*B3S!C89E9E8HY?65=!<:![J?:?9=E5E9=!0?=?9M9G?5!F<!*R!6<8<>?=?5<!*B3S!<!?*B3S!C89E9E8HY?65=!<:![J?:?9=E5E9=!0?=?9M9G?5!F<!*R!6<8<>?=?5<!*B3S!<!?*B3S!C89E9E8HY?65=!<:![J?:?9=E5E9=!0?=?9M9G?5!F<!*R!6<8<>?=?5<!*B3S!<!?*B3S!C89E9E8HY?65=!<:![J?:?9=E5E9=!
5D5<8HP?9=!M?:?E5F9=!C98!=56589=< 5D5<8HP?9=!M?:?E5F9=!C98!=56589=< 5D5<8HP?9=!M?:?E5F9=!C98!=56589=<
5D5<8HP?9=!M?:?E5F9=!C98!=56589=<RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR ptRRRRRRRRRRRRRRRRptptpt
tRrRS tRrRS tRrRS
tRrRS ,DG<D758?5!<>9MJE?>5!5CM?65F5!l!M?D75G<:!?*B3S,DG<D758?5!<>9MJE?>5!5CM?65F5!l!M?D75G<:!?*B3S,DG<D758?5!<>9MJE?>5!5CM?65F5!l!M?D75G<:!?*B3S,DG<D758?5!<>9MJE?>5!5CM?65F5!l!M?D75G<:!?*B3SRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR UTSUTSUTSUTS
tRrRn tRrRn tRrRn
tRrRn %DZM?=<!F9!E85D=C98E5F98!%GEUC!D5!M?D75G<:!?*B3S!<>9MJQF5%DZM?=<!F9!E85D=C98E5F98!%GEUC!D5!M?D75G<:!?*B3S!<>9MJQF5%DZM?=<!F9!E85D=C98E5F98!%GEUC!D5!M?D75G<:!?*B3S!<>9MJQF5%DZM?=<!F9!E85D=C98E5F98!%GEUC!D5!M?D75G<:!?*B3S!<>9MJQF5RRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRRR UTtUTtUTtUTt
tRrRr tRrRr tRrRr
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