• Nenhum resultado encontrado

Dinâmica folicular ovariana e bases para a aplicação das biotécnicas da reprodução em bovinos

N/A
N/A
Protected

Academic year: 2017

Share "Dinâmica folicular ovariana e bases para a aplicação das biotécnicas da reprodução em bovinos"

Copied!
27
0
0

Texto

(1)

!"#$"#%"&$'# () !"*!* (* +*,(! * *-#*./0*#1"*( *234'"# ("!("33'( &$5"46*!-(* +* !'(789*(" :*4#*!"

("!(" ;<<=

(2)

>

!"#$"#%"&$'# ( ) !"*!* (* +*,(! * *-#*./0*#1"*( *234'"# ("!("33'( &$5"46*!-(* +*

?! ! (&$7'"&$ *4 !'(789*(" :*4#*!" #(+5*#789##99 !*#!/#6*@*(

("!(" ;<<=

(3)

;

A%? B

CC

*5"#3*(" #9

* D4*!8*!"-* ##''*!&$#*(6! *!# '"&$4-* #E*(" #*5"#9F("!("7G#9 9H3;<<=

! !"#$I!F*!* (* +*JF *-#* #(""*#(3 !"*!* (* +*,(! *3("!("3 ;<<=

!'(7 :*4#*!"

>9-* C'"&$

-#C!-7*(! 5*#'"&$K* D4*!8*!"-* K (4 (#4 *#

(4)

L

%3 9 * D4*! 8*!" -* ## ' '*!&$#*(6! *!#'"&$4-* #9("!("3;<<=9 ! !"#$ !"# 5"&$ I*!* (* +*3 ? ! (&$7 '"&$ *49F !"*!* (* +*,(! *34'"# ("!("3 *-#*

#(""*#(./0*#1"*( *29

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

-#C!-7 *(! 5*# '"&$K * D4*! 8*!" -* 3((4 (#4 *#9

(5)

T

%3 9 -* 8*!" * 4*!# ## ( ( '*!(* 8('"!(*-*(! *1"#* -* #9("!("3;<<=9 ! !"#$ !"# 5"&$ I*!* (* +*3 ? ! (&$7 '"&$ *49 F !" *!* (* +* ,(! *34'"#("!("3 *-#*

#(""*#(./0*#1"*( *29

( #( :# Q* # !*- * 5 '#*(* * UU* !(( * 59 *# !" (: *# ! #*(* 5 ( '!(*! 8 * ( #*- (! *!-!((* 53"(5(4'#*# '"!(*-88*!* !: 5 (*!*4'-4 (9(( (* 5*- ('"!(*-88*!* !:!4 ( !( 5 (*!*4'-4 ('!##3:@'*(* 5(8"'"!(*-'( (* 8 ( *4#9 *(! *1"# ''* ( '"!(* *# ( @' ( '"!(*- '( (* 8 5( ! 4*! (" 9 ( 4#( *4'( ( '"!(*- *(! *1"# ( 4: '"!(* 8 4: ( #89 *88 ( 4 ((4 (# ! #*(* 5 ( * (8 U*( 4 ':#*5: * ( !*- ( #* 5#9 UV# ( * !# ( '"!(*- (# 8 !(( ("5 'CV (*8*!* * #4* (* * ( 84# * '"!(* 3 U*("( ( 8 #("# (!(* 9 4: '"!(* *4# ( *#(*"( ( 5 (*!# 8 4# 84# 5 (*!: *4'- : ( #8* 5 (* 4:# ( 5 (*!: * 8* 84#9 "'-"(* ((4 (# #*5 ( '- ( ( 4! *#4 8 4* !3 U*! #"(# * 4 : 8*!# -"(* 5 #*4"( "#:84:'"!(* 9(4:'"!(* 3 ( (! *1" 8 -* '" !(" 5"* : ( #-5* "(#" U# ( #'*( "4 8 1"*(: !:(#9 ''*!(* 8 *(! *# # V U58(84 -* '"!(*-':#*5: V U58 (*!(* 5* *88 (# -* 4 (#9

W:U#7'"!(*-*(! *1"#3-* 8*!": 4*!#34 ((4 (#9

(6)

X

"4+*

>9 9999999999999999999999999999999999999999999999999999999999999999999999999999999> ;9 %? 99999999999999999999999999999999999999999999999999999999; ;9>9>9 * D4*!8*!"-* 9999999999999999999999999999999999999999999999999999999999999; ;9>9;9 (*!"*#!*!#(#8N4# Y ;9;9 ##''*!&$#*(6! *!#'"&$999999999999999999999Z ;9;9>9 ##''*!&$* #4* &$(*8*!*4(4'8*@99999Z ;9;9;9 ##''*!&$( #8N !*4*P#99999999999999999999999>; ;9;9L9 ##''*!&$#"'-"&$99999999999999999999999999999999999999999>L ;9;9T9 ##''*!&$#'*&$8*!"9999999999999999999999999999999999>T L9 999999999999999999999999999999999999999999999999999999999999999999999999999999999>X T9 [%? 99999999999999999999999999999999999999999999>Y

(7)

Y

>9 ("&$9

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

# --*4 ( # ("*# (6! *!# #S 8* '##M- !4 * ( # #8& 4 ( 8*#*5* '"(*- # *4*#3 #(* ! #"# 4! *#4# !4'@# !4 ### 54 * ( *8 (# ! *&P# 4* (*# #*8 (#&#-* #9

(#(! (@(3##-*#$(4R(*-8Q"4! @$ ( 8*#*5* '"(*- -* 8N4 -* # *(6! *!# '*!# '"&$ 4*# *8" *# ("4 (3 8*4 "!* # 4! *#4#8*#*S5*!# #1"*###*4###(! 5*#9

(8)

\

;9-*#$**5+8*!9

;9>9* D4*! *!"-*

9

-* 6 "4 # *4*# 46#(*!# 1" (N4 (*!* 4 ( #" 8*#*5* '"(*- 4*# 4 #(" '8" 9 # 0(*4# 6!# !4 - ( "(## 58* ( #( ' -*3 ( "(#3 # #("("# -* # "(* #3 '4*(*" * 4*# ( *4 ( # ! (!*4 (# 1" !4 # -+*# 0( # 8N4#3 5 !*! #( 5-*9

# 8N4# -* # #$ ! #*# '*6#(*!# $C#(!* *#3 '# ( !*! #( 4 * (-# 5"# '@*44 ( ;> *#9 " ( !*! ! "4 !* - (# 1" # '(4 # !*!# #"#1" (# (6 1" "(S*# #R *4'* ' "4 5#(&$9 # -+*#3 5O # 84* * #3 #$ #' #+-*# ' '"&$ 4("&$ # !6"# 54* (*-#84* * #I#S!*(#J31"## --4 * (*#8M!"#3 84# * -* 8( *49 '!## 8*!"5N # I!#!*4 ( 4("&$ 8*!"J (4 * M!* -* 8(3 # 1" ' # "4 4* * ### 8M!"# -* !4'( #" 4("&$ -"9 4**3 " (#"!#!*4 (-*5 '!##!4(#*8*!"9

(9)

Z

I!((:!"#3>=ZYJ948M!"#'-(#(+5*84&$ (( 4'( !*#* ((*Q(##'* !*'*#4O *##(S*# -* #3 # 8Q4 4 1" (*# #( ( '1" # 3 '( (3 # --*4 ( 8*!" '!! ! * ' ( #(S*# S5 # I!V:!"#3>==YJ9

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

(10)

=

4* D !*I*-!"#3;<<>J9##44 (6'4* (@'##$ !'(#'4O *"(* *Q (IAJ !4!6"#5 "#3 '## 8M!"#'5##*-4 (4*##' #*-&$A9

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

(11)

><

5"&$ * D4*! -* !*! #( # + ' "4 4! *#4 " S!* 1" -- *'(+43 *'S8*# -+*#9 *'S8*# (*#!("#5 (8* #3 AA9#M (##!&$ 4##(+#! ('*4+*% A*'(D4*!9 A6#!( 4 * ! #(*("(*-3 *#( 63 4* '( 4O * 6 * 4#4 -!* !4 1" 6 '"Q*9 /+ A 6 #!( 84 '"#+(*3 *4*(4 ( 5" ' *&$ '"#+(* % A !*!"&$ '( IV "44* #3 >=Z;J9 #!&$ % A 6 5" '# #(S*# 5 *#3'5#( #(*31"! (4#!!(M#(*!####!&$ 5!*!#(9

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

(12)

>>

* #"8*!* (# ' * "Q* "4 ( '*! A3 1" *4' 1" ! -"&$9 '!## 1" ! # !"(4 ( "4 ! R" ( 8M!"#4* M!*# --*4 ((6(#*8M!"4* (3"#" -"&$36! !*!4 !#!*4 (8*!"9#(8 O4 ! "#1"(-Q# (* (-"4!*!#( -!9

;9;9(*!"*#!*!#(#8N4#

-* #** 6 !4'#( ' Z< _ *4*# 3'#"-* (#"'**1" 4 (*#44* (#('*!*# " #"('*!*#9 # ! *&P# 4* (*# (# (4'("# "4*3 !('#*(# 85*# *@ 1"* #$ #' #+-*# '

'4* D !* # #*9

5 6 *#(* ( 5 " 4 4"*(# !!(M#(*!#9 " 8*#*5* '"(*- * D4*! 8*!" -* (464 *8444"*(#' (#!4'##" 9( (3#$#(!*# #("#8!#4 ( #'(*!"*#### *4*#9

45# *4*#Q"M #*84#("* # 1"! ! *D4( 4+@*4 # #("("# -* #9 N4# Q"M # '# (4 4 *D4(4+@*48M!"4* (I><C>;44J!'0(I>\C;>44K S! "#3 >==LK *5"*! "#3 >==\J 1" 8N4# ("* # I>TC;<44 ;<C L<443#'!(*-4 (KS!"#3>==LJ9

"(# (# 4#(4 1" 1" (* '5#( * '8*4 # 1" #" 364#44 ## M-*##6*!#'5#( # 1" #" I5# !"#3 >=ZTJ9 I>=ZTJ#"5*"1""44 #Q"M #6#"("4 4 #'#(#(*4 '6C-"(S*A9

(13)

>;

#-!#" 9 (( (-*8*!4"44** !*N !*1"( # !#!*4 (8*!" (!*!#( I>Y_J!4' " I<_3 a<3<>J9 *!# 1"( # (464 84 ! (# 4 -*# 4 IY3Z_K #! "#3 >==XJ -!# I=3L_K ,*("4! "#3 >==YJ9 ! (+* 1" 8* #- ' "4 @'*4 ( !4 -!# 4 I-Q!"#3;<<<J9

4&$)''"&$8*!"#-+*#3 -*# 84 (#!4! ( 4*#8M!"#!44 #X44*D4(4#"# -+*#1" -*#" I5# !"#3>=ZTJ9A+(464-*N !*# *8 (# !4'(4 (# &$ # -+*# !4 8( !#!*4 ( #4 ()* #"* % 94#("8*(!4 -*# I"(* *!"#3 ;<<<J ((4 ( !4 #4((8* IJ "4 (" # ! ! (&P# '#4+(*!# % C 04 8M!"# '1" # IaX44J 84 #4 (1"8*#- 5" I% 5!"#3>==>J9 (( ( (@"4 ( 048M!"#8*4 1"1"8*( # " 9

"(##("#((44 *8#(&$!*-"&$4 9 "&$ !* 8* ( !4 4 1" " 9 I 3>=\YK%* ("3>==<J9"&$4 *8#(&$!* # 6'@*44 (><#3!4-*&$ (>3L;<#I%* ("3>==<K* *!"#3>==ZJ9#!!(M#(*!#* (- ( '6C-"(S* A -"&$ (464 84 #("#9 # ! ! (&P# #6*!# A " ( '6C-"(S* A 84 4 # 4 -!# 4 1"4-!#*(D *!#I" K 3>=\YJ9* (- ( "4 ( #6*! #(* (6 '*! A 8* #*5 *8*!(*-4 ( 4 4 -*# 4 1" # 4 @ A8 " # A8 I " K #!"#3>=\ZJ94 (3 I>=\YJ-*8*!"1"8N4#4 '# (-4 '6C-"(S*A4*#! !*!#(-"4 (# I>Z3X b L3>3 '*# '6C-"(S* AJ 4 !4'&$ )# 8N4#A8I;L3Lb;3>J"!"Q4 (4 @A8I;;3;b;3YJ9

(14)

>L

;9L9##''*!&$#*(6! *!#'"&$9

# *(6! *!# '*!# ) '"&$ *4 84 #' #+-*# ' "45 "4 (1" (*((*-1"*((*- -* !* 9 *!" -* ( "(** ### 84 (# (! S5*!#3 1" "(**Q# !4 !*(6*#1"#9!(4'3 (+-*#- &#84! &# 8* 4 ( # '!*4 (# * "&$ -"&$3 #* ! *Q&$ -"&$3 '"&$ 4*P# !*'#-&$ 4*P#9#(6! *!# $C* -#*-#"(## 58*'#'*&$S!*(# *4*#"(#'6C'0#3#(6! *!# $!*05*!#'( #8N !* 4*P#(* (N4!# --*4 (#(! 5*#'*!#) '"&$9##!5( (*-#'+(*!#6# ! !*4 ( 8*#*5*#4*P#38*#*5*'"(*-#8N4#-* ##"# #'#(#(4* #* (8N !*#@( #'-!#'449

;9L9>9## ' '*!&$ * #4* &$ (*8*!* 4

(4'8*@9

(15)

>T

!* ' # * 4* 4 # 3 !"R !4'(4 ( '"(* '# ( '(*!"*# F !* !"( "&$ !4 -'! ("4 *8#(&$" ('M *(I* *!"#3>==ZJ9

B" ( #( 'S#C'(3 "*QC(Q *-* 5 I>===J -*8*!4 1" 8N4# Q"M # !( (# !*# '#( #(!4 !*!*!* -* #4 ( ( ;>\ ;\Z *# 'S#C'(3 '# ( "4 * (- ('(#>\>=4##9

#( ! +* (! 5* * #4* &$ (*8*!* 4 (4' 8*@ #"5*" ' ' ! ### !" # #"'* ### 8#9 # '(!# #* ! *Q&$'* #4* &$(*8*!*4(4'8*@I J"#!4* "Q* 45N !* "4 - !#!*4 ( 8*!"3 ! ( "&$ !#!*4 ( 8*!" (6 #(+5* '6C-"(S*3 #* ! *Q * #&$ (* 8 ( '5#( @S5 I*4' ( "*!" " *#'#*(*-* (-5*#'#*(*-* J S5 *#'#*(*-* "Q*#'#*(*-*-"&$#*#'#*(*-* ! *#'#*(*-*Q4(## *#'#*(*-*4*#'#*(*-*# I"#* ! "#3 ;<<YJ9 ## 84 * "Q !*!*!* 'S#C'(3 "Q* * (- ( '(#3 64 *#' # #-&$ !*3 * #4* (## *4*#R" (#3 "444 ('6C#(!*9

(! 5* ##* "(**Q&$4O *# ("*#" #* (6(*!#38*(##4 (##4O *# S5 #1"! (4!*! #( -!9 # *# 5 # 5"'# ' #* ! *Q&$ -"&$ #$ # '5#(+5 # " '5#( # # '#(5 * #3 1" '4 # ##!*# !4"(##"#(D !*#8*4#(##"(##R#9

(16)

>X

R(*- 6 * ** '5##$ 1"1" !#!*4 ( 8*!" R+ - &3 !4 (#* "4 8M!" 4* ( '# ( '#(* * M!* "4 - !#!*4 ( 8*!" 4 ( 1"( 1"* ( * ((4 (9 # '(!# ((4 (# ("*#3 ' (* 5* ## R(*-3 # '5#( # " '5#(+5 # #$ ##!*# "(# #"#(D !*#9 !4* &$4* *#(&$'5#(+5 "'5#( !4'*!&$ "4 #(S5 * * !#!*4 ( # --*4 ( 8M!" 4* (3 *4'* -"&$3 ' 4! *#4 $!!%"& 5(*- @!* *'(+4*'S8*#! #1d (#"'##$ -"(S*A9@*#( "4#6*#(S5 #*#' M-*#'#(8*49#(*>\e3"4#(S5 &$ !"(3 ##!* "4 8 ( '5#(+5 I 5#(4(J3 8* #' #+- ' * **&$ !#!*4 ( 8M!" 4* ( > * 'S# ((4 ( !4 X45 #(* >\e3 #"( 45N !* "4 !#!*4 (8*!"3446*3T3L*#4*#(IS!"#3>==XJ9 Q( #(*3 "( #(S5 &$ !"(3 '*! # X453 * (4"#!"3;T#'*#* #&$"43#"(" 45N !* "4 - !#!*4 (8*!"X3T*#'*#* M!*((4 ( IS! "#93 >==YJ9 "( (*' #(S5 3 !*'* ( #(* IJ3 8* (#( ' #- #" 8*!+!* #* ! *Q&$ 45N !* "4 8*!" $#4#("8(*-IS!"#93;<<YJ940(*4#(S5 '# "(**Q'#* ! *Q45N !* - 7-(#(*3 # X45 ##!* "4 '5#(+5 I 5#(4(J3 # L45 #"(" 45N !*"4 X3\b<3;*#4*#(IS!"#3;<<YJ9 #-C# !( -* #'#( #R 1" 4* *#(&$ *8 (# #(S5 #3 '64 # #$ ( (*-# *4'( (# 1" # #R 4##"(# #* ! *Q&$45N !*"4 !#!*4 ( 8*!"4-* #9

(17)

>Y

"(5"'##"#(D !*#"(**Q# ! (!*!#( #$#'#(5 * #9#(#8" !* 4!45 (#"(M(*!#4-!#1" #($ !*! 44 (3 (4* ($ 1" # M-*# '5#( 3 ' *# !' 0( '# (9 # +5# #* (6(*!# I!'#( 3 * '#(3 ( "(#J3 #$ 4*# '( (# 1" # '#(5 * # ("*#9

(#'(!#("*#3#'#(5 * ##$"(**Q#4( #6(*4 ** M!*((4 (3'S@*444 ((* 8 ( @S5 '5#( " '5#(+5 9 '*!&$ '#(5 * 3 !"4* "(S*# "4 '##M- !' 0( '# (3 #4)(*8 ('5#( 3'4*("4(*4 (&$'#*(*-''(#(S5 3 *'(+4*'S8*#3!"4* '*!*&$ A! #1" (-"&$9

* "&$-"&$(46468*('4* *#(&$4O *# @S5 #9#4O *#"(**Q#("4 (#$ Q(#(*IJ3 !*'* (#(*IJ"% A9##(S5 #1" 4* *#(# "#N !* "4 !' 0( '# & "4 8M!" 4* (3 ("4 *'(+4*'S8*#84#(*4"*&$% A*'(D4*! A'*'S8*#9 "QC# ($"4'*!*&$A1"6#' #+-' # --*4 ( 8* 8M!" 4* ( ! #1d ( -"&$9 4#4 843 4* *#(&$ @S5 % A (" *(4 ( # *'S8*#* "Q* '*!A-"&$9

(18)

>\

44 ( (* 8 ( '5#( #"(" 4 4* (@ ! !'&$'S# ILZ3=_XX3>_J34-!# '*#I\Xb>=*# 'S#C'(J34 (*#'#(9

;9L9;9##''*!&$( #8N !*4*P#9

! (@( (" !#! ( 44 ( 5 6(*! # # -* #!(*(3( #8N !*4*P#'S#8(**Q&$" #"5*" !4 84 ( *4'( ( ' * ( #*8*! 4@*4*Q *#(*"*&$ 4(* 5 6(*! '- * ( 4!# 8N4# !4'-4 ( #"'*#9 '##*** "4 4#4 8N4 '"Q* * 04 1" (* 4*P# 4 "4 'M 4 1" '* '"Q* ("4 ( #4 ( "4 !*3 "Q* 44 ( (4' !4 1" 5 6(*! # # -"* '#*(*-4 (3 1" 5 "4 5 5 4 '"(*-*9

( #8N !*4*P##*C# "(**Q&$4$#CC"5"3 8N4# "(#3 4 '( !* 5 6(*!3 1" #( -Q*# -$ ! 4*P# 8N4# #"'*# #$ #' #+-*# ' 5#(&$ !*&$ #(#9

4 ! *&$ (6! *! # 8N4# !'(# # 4*P# -4 #( 4#444 (!*!#(41"#($#8N4##9## 84#$!'Q#'-"44* ("(* 1"'!5 4*$9 #(# #$ ( #8*# ' -( #6(*4 * 'S# -"&$ !'(9#(44 (!'('# ("4!'0((*-3'"Q* '5#( "44* ("(* 1"9

(19)

>Z

84 !##+*"45 1" (* *4*#'#("4( 8N4#!*! 44 (1"9

("4 ( !4 (! 5* ( ## 4 R $ 6 4*# !##+*9 # !'(# (N4 #" -"&$ #* ! *Q ## 84 # (4 "4 1" (* #R *4*# ' (# ' ( #8N !* #4 !##* #-&$ !*9 #*!4 (3 #* ! *Q&$ # !'(# #5"#4#4#'(!## --*#' 3*8* " *!4 ( 1" # *4*# $ #$ * #4* # 44 ( -"&$ !4 # 4*P#!!\*#4*#(9

;9L9L9##''*!&$#"'-"&$

9

( #8N !* 4*P# 8(**Q# !"(*-# (4 #* 4'4 ("(**Q #0(*4#6!#9(6! *!* *!*##* '*!&$ 4O *# 1" #(*4"##4 !#!*4 (3 # --*4 ( -"&$ "4 04 5 8M!"# #*4"( 4 (3 * ** 4! *#4 4* D !*8*!"31"*4'**-"&$4*#"48M!"9#*C # (464 * #4* &$ 'S# #-&$ !*3 1" #*4 ( ( 4 #8*!* (9##! (@(#((4 (#'* #4* &$4(4'8*@ 84'(##((4 (##"'-"(S*#3#!( !##* (!(!*(464 ##(6! *!9

(20)

>=

("&$ # !'(# ' A # 8M!"#9 ## 843 (4C# # --*4 ( 40(*'# 8M!"# -"(S*#3 *4'* 4! *#4 4* D !*31"#*#' #+-'(#*#8M!"#4# --*4 (9

4 - ( 3 '*! # '(!# ((4 ( #"'-"(S*#3'##"C#* *!*'(!!4'*!&$"48 ( '5#( 3 #4 4* *#(&$ "4 #(S5 3 !4 6 8*( # '(!# ' 3 ##*4 8Q 5##$ "4 8M!" 4* ( '4- * M!* "4 - !#!*4 ( 8*!"9 4#4 843 8*4 ((4 ( "4 '#(5 * 6 '*! ' '-! "(S*#"4#(S5 "% A64* *#(8*4* "Q*'*!A 1" #+ #' #+- ' # --*4 ( 8* # 8M!"# ! #1d ( -"&$9###'(!#8!**(44 R#4*P#*5* (@#4*P#-*+-*#'4 #($#1" ()#(*#'S#"(**Q&$ "(#((4 (##"'-"(S*#31"1"4(!&$!*I"#* !"#3;<<YJ9

;9L9T9##''*!&$#'*&$8*!"

9

(! +*#*(! 5*#'*!#'"&$-* #3 '"&$ 4*P# (4 # #(!9 '8" 4 ( # ! !*4 (# # # --*4 ( 4* +* #" 8# * *!* # --*4 ( (6! *!# 8(*-# 4("&$ 8(**Q&$ S!*(# ( " '##M- *8"#$ '*!&$ 4 5 #! ### (6! *!# (S*9 64 *##3 46( '" &$ -* ( #-5* 5"* ' "(##43 ' #'*&$ S!*(#3 #"5*" !4 46( #5"3 '"! * -#*- 4"*( 8*!Q ' !"'&$ S!*(# 4 4 #( ' 4 *'"&$4(S*9

(21)

;<

#"#!6#'*! (0#8M!"# (*#R(*- S!*(1"#(+ * (* ! 8M!"9 4 ' 8(**Q&$ # S!*(# -4 #( 4 8# # --*4 ( $ 4 (#*3 '!"C# 1" *4 # '# ( * M!* # --*4 (8*!"3841"8M!"# R+4#(+5*- &3!48M!"#'6C-"(S*#"8M!"##"* #3 R+ 4 (#*3 #$ * #R#9 * * M!* 8*!" 4 1" # ! (4* 048M!"##!"'4* 04S!*(#9

("4 ( ((4 (# 4 *# (464 #$ "(**Q# 4 *4*# 1"($#"#-+*##'*#9R(*-6@(4 (!"4* 44 ( #'*&$ !4 4 44 ( !*! #(9 (* "4 8M!" 4* ( " "4 !' 0( # -+*# 6 #R (464 ' * #'&'!#!*4 ( -#8M!"#9

# ((4 (# "(**Q# ! #*#(4 4* *#(&$ "4 8 ( *&$ ! (M " '5#( 3 8*4 '-! (#* 8M!" 4* (364'*!&$'#(5 * #'*#!'0(9* "(#( (*-##$'##M-*#3!44* *#(&$ A31"("## 8M!"# #(*4" #" # --*4 (9 4* *#(&$ ' (464"4 ( 048M!"#!"(# 8*!"3(-6# ("&$ 8M53"4 ( *&$8(!#!*4 (#4 () * #"* I% CJ1"("##8M!"#3#(*4" #"# --*4 ( I"(* *!"#93;<<<J9##843''C#-+*'1"#'###'* "44* 04S!*(#1"*'4 *'"&$9

L9 !"#$9

(22)

;>

4# 5 (*!4 (9 ## (" -4 4" ! +* '!"+* #*94! ('(*35 '(###'#1"*##8*# --*8! ' *4*# " 3 1" *84 4"*( #" 8*#*5* '"(*- -* !4'# # 9 #* 1" #* '# ( 5

'4* D !* 3'( ('*!**#*(! 5*#

'"&$' #! !*4 (## --*#8!##Q"M # # !4 #' 4 ### *4*# )# (! 5*# '*!#3 # 4* (# -##41"# ! (5 '( #**3#! *&P# ( (4'(" 85 # *@ 1"*3 '* !*'4 (9 *## 1" # '#1"*## -N4 ( # 0(*4# # 6 *## 1" -* (4* - &*8"#$#*(! 5*#'*!#'"&$ ##'M#9

T98N !*#**5+8*!#9

/3 9K W K 9/K AA /999 "'-"(* * !(("#* 5%8U:%C4 ! (**#9 *49'9!*9-9LL3'9==C><=9>==L9

,3 K 3 9/K A /93 9K c3 9K A3 9K ]3 9K! "# -* !* !!(*#(*!# "* 5 ( #("#!:!* 5"#34 '!U#* #"('*! -* 4 (9 /9 *49!*9\Z3>;=>C>L<;9;<<<9

3 9K %3 9 9%9 4: ( #8 * !((9

*5 5:3-9XY3'9>TZLC>T=Y9;<<>9

(23)

;;

39K39KB39K39 K3%99 "#84 ((4 (#(*4'-'"!(*-'84 !8 #("#8 !((* ('*!!*4(#9 *49'9!*K-9Z;CZL3'9T\=CTZY3;<<T9

39Kc3AK,39AKc39K%3 9K C/3 /999 4'!( 8*!*N !* '"(*- 4 -* # !(9 *# *4'S#* ( !* '"&$ *4 '*!9 * 3'9>>LC>L;9;<<Y9

]3 9K %3 9K ]3 /9K A3 9K / %39#8-* A8#"'-"(* 4:'"!(* * 8 *8#9*5 5:3-9LX3'9><Y=C><Z;9>==>9

3%9K3%9K39K 39/9@5 "#! ( 88*!"U-45 !:* !((9*5 5:3-9TL3'9L>CT<9>==X9 %9K K , K 9/9 *!" U-45 !8(((4 (#U*(#(* Q( !*C-5* -*!#* ( !((9 >L( ( (* 5## *4 '"!(* 3 : :3 "#(*3-9\3'9;;9>==Y9

3 %9K , 9%K ,3 9 K W3 /9K 3 9/9 * ! *Q!* 45N !* 8*!" : -"!* 4 *4#((#!4'5#(5 #:*8 (##(##(*9 *# *4'S#* ( !* '"&$ *4'*!3 * 3#*3'9\>C ZT9;<<Y9

3 %9K ,3 K 3 9 K 3 K 3 AK 39/9 *!": 4*!#* " 8!(( " '#(" ! *(* # * 5 (* 9 7 !* 5# 8 ( ><( 5## #**'"&$ *43-9;94'% 3'9;;>9>==L9

(24)

;L

-* 8*!" -'4 ( * I J *8#9 *5 5: -9XT3'T;>FTL>9;<<<9

3 9WK ,,3 9/K ]3 9 ( 8 ( 8*! -'4 ( #!(* * #' !((9/" 8 *4!* !3-9T=3 '9LLXCLX<3>==X9

39WK3A33%9Kc39K399 88!( 8 #(*C * ** C8 -* 8*!" 8"* -* 8*!# -* 4 #!(* 9/' (3-9=L3'9Z>F=Y9>==>9

W39/K3/99(4'(* #*'(U 5 ('* #* 5 4 I% AJ "(* *Q* 5 4 IAJ #!(* * -*!(4*QU#9 !* 5:3-9>>>IXJ3'9>\L\C>\L=9

%39K39KA39K3/999-* 8*!": 4*!#* I J9*5 5:3-9T\3'9>TZ=F >X<X9>==\9

5*!"(" 5 *Q(* 3 3 ;<<\3 *#' M- #*( (('7EE8#((9895E3!##4=5#(;<<=9

%39KA3%9A9-*U !(('"!(* * (('*!#9( T9#("#!:!#9 *499#(99XZ3'9Y=\F\<\9>==<9

%%3 /9%K c3 9K ]3 9 88!( 8 !4* ( -* #4(('* -* 8" !(* * *8#78*!"''"(* # '*' 4 #9*9'K-9TX3'9=T>F=T=9>==>9

% 9 ( '"!(* * !(( "88#9 4*57 ( (* 3>==Y9

(25)

;T

4Q:5"#3 (Q:5"# C!*# 8 8!" *(: 5 * 8" !* 5 (* -"(* (9/" 8'"!(* (**(:3-9Z>3'9L=XFT<;9>=Z\9 ,39 K3%9K3K 39/9-* 8*!" : 4*!# * " 8 !((9 *5 5:3 #"4*(( 8'"*!(* 9;<<L9

W3 /9K A3 9%K W3 9K %]3 %99 )) "* 5 #' ( "# "(:#*# * ( !:!*! 5 4#(7 *!4*! 4'5*!-* !9*'3-9XZ9>==Z9

c3W9K3KAA39K3WKA3KA39K 3/K%3KA3W99*88 !#* -* !(*-*(: (U h* U#U*!U4Q:5"#3(Q:5"# C !*#84R5 * 8" !* 5(*-"(* (9/" 8'"!(* (**(:3-9\\3'9>=LF;<X9>=ZY9

K 9K A /9/9K 9K / ]99WK #4' ' *4*# !"Q# !4 9 9 M '6 #449 7 " *$ " !* #** ,(! *3 LL3 '9>;\9 >==Y9

A3 9K 3 9K %3 9K 3 9K 3 39K399#("#-*" (#("#C(C -"(* * (- * !(( I J U*( (" #("# #("# * "! U*( '#(5 * ;` 5#(4( #(* -(3 *5 5:-9T=3'9YY\FYZ>9>==Z9

3 99 A -"(* * 4 3 4 i A8 A8*8#9/9 *49!*9TL3'9L<<9>=\Y9

(26)

;X

3%99K 3/99-'4 (84* ( *!#* (( # ##!*( U*( *!C(*4"(* 5 A4 C' ( #"* C*V %U( !(* * 5(* CT(#9*5:8'"!(* 3-9YX3'9>>;C >>Z3;<<>9

A 3 9K 3 9K A3 9%9 '"!(*- #("*# 8 4 !((9 9 "(* *Q* 5 4 -# * -*!(4*Q * 4 3 4 iA8 A8!U#8U* 5;<45#8#(*C>\`9 *5 5:3-9><3'9T;=FTL\9>=\Z9

A3 9KgA3%K]3W9]9 *4 (4'88!(# -* 8*!" : 4*!# * 4 *8#9 *49 '9 !*9 -9LZ3 '9;YXF;\\9>==X9

,C3 ,9K C%3 9 -* 8*!" : 4*!# * #"!VQ"I J!U#4 *(:(*4"(# 5':9 *49 '9!*K-9XT3'9;>>F;;<9>===9

%3 9K A3 9K c3 9]K 3 9K %,3]99 -* "(* 4'5: 8" !(* * 5"# 4 !U#9 /9 *49!*9X=3><;YF><TY9>=ZT9

]3 K3]99*5*!8" !(* # !'(* * 5!(*-*(*# 81"* !* *!5 ('* #9/" 8'"!(* (**(:3-9Y;3 '9X;\CXLY9>=Z>9

- ], 3 % 9/9 '5*! !!(*Q(* 8 -* '*4* 8*!# (* -* 4 ( * '3 -9XX3 '9><<LC ><>>3>==Y9

(27)

;Y

Referências

Documentos relacionados

Após uma dose única de 3 mg ou múltiplas doses de 6 mg duas vezes ao dia, o clearance (depuração) médio oral da rivastigmina foi aproximadamente 60-65% mais baixo em pacientes

Esta pesquisa teve como objetivos pesquisar como estão sendo utilizadas as Tecnologias da Informação e Comunicação (TIC) nas escolas que atendem os Anos Iniciais

Com o decorrer do tempo, a criança começa a ter um controle maior de seu corpo, e ao ser estimulada corretamente, seus traçados ganham formas mais precisas e no estágio de

As investigações pautaram-se no funcionamento do processo produtivo da construção considerando a forma de produção por encomenda, com o uso do trabalho do escravo e do

(UFBA), da Universidade do Estado da Bahia (UNEB) e da Universidade Estadual de Feira de Santana (UEFS), assim como na base de dados Scielo, que tivessem as palavras chave ‘São

Esses achados entram em consonância com as ações de prevenção, promoção e tratamento mais frequentes do Ministério da Saúde, são estas: incentivo ao aleitamento materno,

A ética social cristã exige um modelo de compreensão que apresente a racionalidade da praxis e a teoria da acção colectiva uma vez que o enunciado da fé se deve

Outra alternativa são os modelos baseados em aprendizagem de máquina (AM), representados por metodologias tais como Redes Neurais (RNA), Árvores de Decisão (AD) e seus