Densidade e caracterização de bactérias diazotróficas associativas oriundas de diferentes sistemas de uso da terra na Região amazônica

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; * =2 \E ( ?NN;>( " * E $+ $ 7 * # T ? 4 1 ? A =0 \ ( 344?>( " + =) > H 7 * $# ? 4 ? A 1 A 4 E =R ^ ] \ 344A>( 0 7 + $ + # B E # = W2>( 6 ( =344<> + " ! ; = W2 > T # * B $ + W2 * # " # # 7 # E + # T ( 0 7 ( ! + # 7 =- D > G * 7 8 =2 E ( 344@>( " $+ E * BU $+ E =2 ( 3444>( 0 7 + * 8 7 = . > # ( ) $ 7 " $ * $+ B * *7 =- Z ( 344@>(

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M 9> =0)%# ) 7 7 : " =2 ( ?NM5>_ " =2 ( ?NN5>_ " =6 EE * ( 344?>_ " =, ( 344<>_ " =. ( 344@>_ " " " = ^ ] \ 344@>( 7 $ # # / ( 7 " " *$ # # # =2 ( ?NN5>( " 7 " * . # ? < =2 ( ?NNM>( - [ ( =3443> #7 " ( 7 *$ + + S E + =6 Z ( 344@>( + 7 * ?N A@V B =6 Z ( 344@>( 7 " * =. ( 344@>( 9? =0)%# ) # # / * # * W2 ( ) * ] # E ( =?NN3> 7 & * * ( @A 7 E = 3445> * *7 " =/ ( ?NNA> " =0 ! (

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?4 * # $ =2 ( 344A>( 0 8 B E * " ( 0 7 BU " ?5! =NM ?44V> $ E # " " : " _ " _ " _ " _ " _ " _ " _ " " =2 ( 344A>( " #7 ( -( =344?> * B " && -? E + <5 @MV( 0 7 * A N 5 <V( ( =344?> * B " ! )D,3 #7 * G ( ) * + = * G * > X ( ) * ! )D,3 # + * * *S * + # * G ( " * # " C # E ( =?NNA> * 7 E # * " W 6( " # E * + B * # $ + * E +( ) " B E # + + * + E # + W 6( 9 + * ( 7 B $ E * #

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?3 # # B $ 7 & ( ^ E =344<> # H *$ + * + * & ( ) & # * # # G * " * ( ) 7 * $ 7 ( 0 7 B # 7 * + G + 7 $+ + =/Z E[ ( 344?>( C #7 7 E * T =/ ^ / ?N;M & Z ( ?NNA>( C E ( =3443> 7 + # + * + + - - - B " ?5! B B B N;V ( 0 7 #7 E =& . Z ^ 6 ?NNA>( 9@ 7* !# +# - -!)7 +) 6-# +# -#"# -#1%) #- 7 %#%$ 0 -7#-7 B $# X $ + S 7 B $# # 7 ( ) ( & ( =344@> # 7 + * *$

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?< # 7 # ( * # # * : $ E # # 7 # ( + * * 7 S * + H" # = (> " # = (> = > + + # ( ) * B * + * B * # # $ _ 7 E * $ # ( " # ( =344@> * $ /" * X $ 7 ! ! /0 ;3 # # # " # H" # ( ) * $ # 7 7 ( W * * 7 + * ( ) H" # B * " ( #7 7 ( 0 # # E * $ 7 B * + # 7 7 + ( * ( 7 X

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?@ * $ * + ( 9A -!6+#- +) +)0- + +) ) + ,)%- + +) B * + * B + * 7 S ="& > + =& E ^ `# ?NM@_ ( 344?_ ^ & 3443_ & ( 344@_ 2 ( 344A>( ) + + * # =& E ?NM<> /. H" # " # = `# ( ?NNA>( /. " # H" # * 7 " ( ( # E 7 7 =& E ^ `# ?NM@_ ( 344?_ C E ( 3443_ " # ( 344@>( ^ & =3443> * B * /.( ) + : * 7 "& + * ?< 7 7 _ " # + *7 + * + * ?3 * 7 ?4 7 B $ 7 ( ) * B # * + *7 7 B * " ( " *

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?A * B " * BU B /.( ) E * $ 7 T + * * # ( . T # * 7 ( 7 $ * =! ! /0> * $ # 7 + * ( " # ( =344@> 7 7 ! ! /0 # # B * T E ( ( =344A> * 7 7 +8 ( 0 7 * B E * ( 7 + * # B $ ( + $ # 7 + * 7 * S E # * ( ) B B * # -" # =?5! 3<! A!> ( ) -" # 8 7 * T $* # + * * # ?5! + T # 7 =c ?NN?>( ) ?5! * = D->

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?5 B BU =c # ( ?NN?>( 7 * * 7 ( BU # # # # / 2 \ BU G E ( <;3N@ ?@M55< 7 BU " E # / 2 \ =C Z 3445>(

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?; < <9; *) !# $)% -0 # E * + G % , - %. / & = "0 >' 2 D & * d . 7 6 &7 ( " 2 # E * ( ( & ! + & ( + +8 $ $ 2 G D + ( + / # .. " 2 G D ?(?44 \ & # - ! T = ?>( $ E 33 * $ C D ( 0 + T B * ( ) G 7 # # # + * ( E =! C> + G 7 + * + B # G (

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?M ./ - ? + = 2 G D $ " / # .. $ 2 G D >( <9 7#-!% $)7 +) -#"# # $ G * + NM ?44 A4 = ( 344A>( * 344@ * ?3 * 4 34 ( 6 0 1 % #, " 04 )0B 7 0 #0-! 0!

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3? > =& E ( ?NM<> = 3 >( ) * ?@ X 3;eD( + * * 7 7 S ="& > + %& # # " # 0 ' =& "0!> =c ( ?NMM>( C B $ 7 * $ * ( ) B * $ * ( "& * $ B *$ B @ # *$ ( * * B $ B * # 7 B $ * * ( " * * =H" # " # /. >( D 8 * * * $ ( ) B * $ * /" # * + ( ) # * + 1 34eD( <9> % !)% 3 4.# )7 7) # +) 6"! ,# ) 1 ! ! + * + /" # = 3 > $ 3;eD( $ * _ X 7 = >_ * = * >_ = \ # # >_ # = # * >_ *$ = >_ = # >_ = B # $ >_ E = S # E > =# >(

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33 T * $ * : 2-??44?C = >_ 2-???@4C ₌ _{>_ 2-??<@4}C = >_ 2-??4M4C ₌ _{>_ 2-???;A}C = > 2-N44@ = ( # &( !( & &/>( <9? , " 4.# + 7#%'#"#$ )"6" % )7 7 %#- 5* # 5! # ) !)-!) +) % 7 ) * B * 7 * =) Z 2R@4>( 3;eD $B # * X # * ( $ # * _ * 7 _ # X E # = W2>( ) * B / 2 E [ [ =] ( ?NN<>( * * /" 3;eD( 8 * * _ + * # X ( * # ? # ? NAV <4 # * ?4 ( 7 * # #G =" \ R .. >( 7 / * + / E (

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3< <9 0&" -) *%#!/ !% ,/- +) )")!%#'#%)-) )7 $)" +) *#" % " 7 + C F ( D ) * + C] =% Z Z '> C] $B B # @ 3MeD # ?34 = < >( ) * * M4 R ?44 ?4 @eD # % ' 7 B * + " 2!( * ;4 7 # % * ' 4 N =CC > 4 ? ! ! 34V( * B # E ?4 NAeD * M4 R ?44 ?4 @eD( * # * = /0> 7 =?N;4> * H \ =?NMA>( <4 $ ?3V A V ( # * # $ 4 ? ( # * E * * H =!G> B !Gf 9 g # g ( ) * 7 /& = \ > * ="C!]! 3(? >( <9@ ! , + +) + 0 !%#$)0 -) !% ,/- + %)+64.# +# )! ")0# C D 7 * + [ E =?N55>( * * ?4 =A > T 3MeD 7 * $ $ ( * * $

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3@ $ # * ( " =% Z Z' - > * + Z 6 7 # =D D3> _ ? * G # ( ? E * ? # * 7 * =, ! <@44 %>( <9A 7*" ' 4.# ) -)G6)0 7)0!# * % " +# $)0) ; +# % 1#--#7 " ) * /" ( D 8 * # % * ' ?444 7 B ?4 NAeD( 0 * $ ?4 + $B 34 * + = D-> * ?44 ( * * 4 3 & $ 3; =A / /CCC/ CDDC//DCD /> ?@N3-=//CC DDCC/CC D/ DCC> 3 A & 7 ?R D- 4 3 & "C 4 43 C B " = C& C B " . >( * * + 0 * & Z h _{E (} _* * N@eD A ( <4

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3; ) E * $ ( " " * + *7 ?< 7 B ( 0 /. * # " 7 + * ( " * BU " + *7 g + * + * 7 ) E * /.( ) # $ B (

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( C #$; /" 6" -$% 7 +) -# "# F; D 4 ? 3 < @ A ./ - < "S ="& > 7 " # <4 * =! C> +8 ( 2 * NAV( 3M

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3N

E;F;E;F @E;F ?E<F>E<F>;E>F E;F ;E>F?<E?F AE?F@?EF ?EF;@_EF;@ E?F E?F@EF @EF ;E<F<<E>F?AE?F@E?F@AE<F<@E>F EFHEFAEFA>EFAEFH;EFHEFH>

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<? ) 7 # 7 + * " # =! C> * 5( 0 " # E ! C * - D " 7 * $ ( " ; /.( ) # : _ - D , E * D " ( 0 E ! C D , E = M>( 0 /. E * $ ! C( "#%)-! 9 #, $%#'"#%)-! #4 9 )" ( -! $)7 ( C #$; /" 6" -$% 7 +) -# "# F; D ? 3 < @ A ./ - 5 &7 =! C> +8 S ="& > 7 + * " #( 2 * NAV(

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(80)

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