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SOCIETY OF VERTEBRATE PALEONTOLOGY
NOVEMBER 2014
ABSTRACTS OF PAPERS
74
th
ANNUAL MEETING
Estrel Berlin
Berlin, Germany
November 5 – 8, 2014
HOST COMMITTEE
Nadia Fröbisch; Jörg Fröbisch; Oliver Hampe; Heinrich Mallison; Johannes Müller;
Juliane Röhner; Daniela Schwarz; Florian Witzmann
EXECUTIVE COMMITTEE
Cathy Forster, President; John Long, Vice President; Philip Currie, Past-President;
Glenn Storrs, Secretary; Ted Vlamis, Treasurer; Elizabeth Hadly, Member-at-Large;
Christian Sidor, Member-at-Large; Paul Barrett, Member-at-Large
SYMPOSIUM CONVENORS
David W. Bapst; Andreas Christian; Jussi T. Eronen; John R. Hutchinson; Martin
Kundrat; Michelle Lawing; Graeme Lloyd; Nicholas J. Matzke; Andrew Milner; P.
David Polly; Daniela Schwarz-Wings; Eric Snively; Jean Sébastien Steyer; Florian
Witzmann; April Wright
PROGRAM COMMITTEE
Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Heather Ahrens; Brian Beatty;
Chris Brochu; Richard Butler; Ted Daeschler; David Evans; David Fox; Pat Holroyd;
Marc Jones; Christian Kammerer; Matt Lamanna; Erin Maxwell; Josh Miller; Jessica
Miller-Camp; Kevin Padian; Lauren Sallan; William Sanders; Bruce Shockey; Mary
Silcox; Michelle Stocker; Rebecca Terry; Paul Upchurch; Aaron Wood
EDITORS
Erin Maxwell; Jessica Miller-Camp
Designed for palaeontology courses in biology and geology departments, this leading text will also be of interest to enthusiasts who want to experience the flavour of how the research is done. The book is strongly phylogenetic, and this makes it a source of current data on vertebrate evolution.
Study aids include key questions, research to be done and recommendations of further reading and websites.
Read sample content and order at www.wiley.com/go/benton-svp Save 20%! To benefit from this exclusive discount make sure you quote
GESAMTÜBERSICHT
GENERAL PLAN
Fo ye r 1 G al er ie (1-4) / G all er y (1-4) VI P-G al er ie / VIP G al ler y Kü ns tle rga rde ro be (m eh rst öc ki g) / Ar tis t D re ss in g R oo m (mu lt is to rey ) Innenhof (1+2) / Inner Courty ar d (1+2) Foy er 3 ECC - Raum 4 (2. Etage) / Room 4 (2nd Floor) ECC - Raum 5 (2. Etage) / Room 5 (2nd Floor) Zu fah rt / Ent ran ce C on ven tio n C en ter Zu lie fer zo ne / D eli ve ry Z on e C on ven tio n C en ter Zu gan g R ot un de / En tr an ce R ot un da N eb en ei ng an g R äu m e 1 -3 / Si de e nt ran ce s r oo m s 1 -3 42 43 44 45 46 47 48 49 50 51 52 53 Ko nf er en zr au m A nt ib es / Co nf ere nc e R oo m An tib es Bu si ne ss C en te r / Bu si ne ss C en ter Es tr el St ub e G ep äc kr au m / Lu gg ag e R oom Re ze pt io n / Re cep tio n Co nc ie rg e / T ic ke tc ou nt er Ko nf er en z-Ser vic e / Co nf er en ce S er vic e G el dau to m at / Ca sh M ac hi ne Te le fo n / Te le phon e Ko nf er en zr au m S tr aß bu rg / Co nf er en ce R oo m S tr aß bu rg M in im ar kt / H ot el S hop Kl ein e G al er ie / Sm all G all er y Ko nf er en zr au m L yo n / Co nf er en ce R oo m L yo n G ro ße G al er ie / Lar ge G al le ry Ko nf er en zr au m P ar is / Co nf er en ce R oo m P ar is Es tr el S aa l A , B , C / Es tr el H al l A , B , C Fo ye r E st re l S aa l / Fo ye r E st re l H al l Pa ss ag e E st re l S aa l / Pa ss ag e E st re l H al l Su n-Th ai - A si at is ch es R es ta ur an t / A si an R es tau rant Po rt of in o - It ali en is ch es R es ta ur an t / It al ian R es tau rant La P at is se rie San ss ou ci - In te rn at io na le S pe zi ali tä te n / In te rna ti ona l S pe ci al iti es Pi an ob ar At riu m ba r O ran ge rie ( 1. E ta ge / 1st F loo r ) Te rr as se / Te rr ac e Ko nf er en zr au m S ai nt T ro pe z / Co nf er en ce S ai nt T ro pe z Ko nf er en zr au m Ni zz a / Co nf er en ce R oo m Ni zz a Ko nf er en zr au m C an ne s / Co nf er en ce R oo m C an ne s Zu fah rt H ot el -T ie fg ar ag e / En tr an ce U nd er gr ou nd G ar ag e H ot el Ta xi st an d / Ta xi R an k So mm er gar te n / Su mm er G ar de n Bo ot san le ge r / Lan di ng P ier Bah nh of / Tr ai n S ta tio n At riu m Ro tu nd e / Ro tun da (Ü be rg an g z um F es ti va l- u nd C on ve nt io n C en te r / Pa ss ag e t o F es ti va l a nd C on ve nt io n C en te r ) W eit er hi n f in de n S ie i n / In a dd iti on y ou f in d o n: Fl üg el 1 / W in g 1 1. E ta ge / 1s t F lo or : - S au na , F it ne ss , M as sa ge Fl üg el 2 / W in g 2 1. U nt er ge sc ho ss / Ba sem en t: - E st re l S m ok er s L ou ng e & B ar - T ax is tan d / Ta xi R an k - T oi le tt en / To ile ts 1. E ta ge / 1s t F lo or : - E is w ür fe lm as ch in e / Ic e C ub e M ac hi ne Fl üg el 3 / W in g 3 2. E ta ge / 2n d F lo or : - E is w ür fe lm as ch in e / Ic e C ub e M ac hi ne Fl üg el 4 / W in g 4 1. E ta ge / 1s t F lo or : - E is w ür fe lm as ch in e / Ic e C ub e M ac hi ne
Estr
el Conv
ention Center
Estrel Hotel
Fl üg el 1 / W in g 1 Fl üg el 2 / W in g 2 Fl üg el 3 / W in g 3 Fl üg el 4 / W in g 4 Re st au ra nt s / G as tr on om ie / G as tr on om y Co nf er en ce Lif t To ile tt en / To ile ts Ra uc her ber ei ch / Smo ki ng A re a Co nv en tio n H al l Fe st iv al C en ter 1. E ta ge / Fi rs t F loo r 2. E ta ge / Se con d F lo or 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 30 31 -Bahn A B C BAR 45 45 51 51 51 51 51 51 51 51 43 43 43 42 44 47 43 46 46 53 11 13 50 34 52 14 32 15 16 17 18 19 22 33 9 12 7 8 10 6 3 4 5 23 24 2 1 26 27 28 20 21 29 M 4 1 Bühne / Stage Festival Center Tor 1 / Gate 1 Tor 2 / Gate 2 Fl üg el 3 / Wi ng 3 Fl üg el 2 / Wi ng 2 Fl üg el 1 / Wi ng 1 Fl üg el 4 / Wi ng 4 Sonnenallee ZiegrastraßeCo
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Sekt io n D / Se ct io n D Sek ti on B / Se ct io n B Se kt io n A / Se ct io n A Se kt io n C / Se ct io n C Es tr el S aal / Es tr el H all Ra um 2 / Ro om 2 (E rd ge sc ho ss / Gr ou nd Fl oo r ) Ra um 3 / Ro om 3 (E rd ge sc ho ss / Gr ou nd Fl oo r) Ra um 1 / Ro om 1 (1 . E ta ge / 1s t F lo or ) EC C 49 48 TAXI 25WELCOME TO BERLIN
It is a great honor and pleasure for the Host Committee to welcome you to the 74
thAnnual
Meeting of the Society of Vertebrate Paleontology in Berlin, Germany. The meeting will take
place at the Estrel Convention Center conveniently situated in the southern part of the city, in
immediate proximity to Berlin’s excellently developed public transit system and the city
highway, allowing you get to around quickly.
This is only the second time that the meeting takes place outside North America, and the first
time it is held in continental Europe. Hosted by the Museum für Naturkunde Berlin, our
Committee consists entirely of staff members from this institution. Since the German
reunification and especially in the last decade, the museum underwent some dramatic changes.
The main parts of the exhibition were completely redone, and by now our classic dinosaur hall
not only displays the largest mounted dinosaur in the world, Brachiosaurus brancai, but also, for
the first time since its acquisition in 1880, the famous Berlin specimen of Archaeopteryx
lithographica. In addition, the museum's East wing, which had been left as a ruin since the allied
bombings from 1945, was entirely rebuilt to house the zoological wet collections, and in 2011
was awarded the renowned Prize for Architecture in Germany. We are very happy and proud to
have the opportunity to showcase the museum's extensive renovations at our Welcome
Reception.
Like probably no other city in the world, Berlin illustrates and reflects the history of the past
century. Be it the flair of the 1920’s, the horrors of Nazi dictatorship and World War II, or the
Cold War and the Wall – they all left an imprint in both the city’s structure and atmosphere that
can still be witnessed today. Berlin is also famous for its arts scene and culture. Just to name but
a few possibilities, there is Museum Island, a UNESCO World Heritage Site showing the
cultural history of mankind from its early beginnings to the verge of modern times. The Jewish
Museum and the New National Gallery are further excellent opportunities, and for those
interested in classical music Berlin’s three opera houses and the world famous Berlin
Philharmonic offer just another wide range of options. Berlin is also one of the greenest capitals
in the world with many large parks and recreation areas. And not to forget the very
international culinary landscape with diverse restaurants, bars, clubbing and nightlife!
Germany has a long research tradition in paleontology, and some of the most famous fossil sites
can be found here. It is our pleasure to offer you field trips for every kind of taste, ranging from
the Paleozoic to (almost) the Present and covering important localities such as the Permian
Bromacker quarry, the Jurassic Solnhofen limestones, and the Eocene Messel Pit World Heritage
Site. We also have a variety of workshops covering a wide range of topics that we are sure will
gain your interest.
The 74
thmeeting of the Society of Vertebrate Paleontology represents a great opportunity for
researchers from all around the world to exchange ideas and further advance our discipline. We
are convinced that you will enjoy the meeting, its fieldtrips and its workshops, and that Berlin’s
historical flavor and vibrant cultural life will make your stay a memorable experience. Welcome
to the city where the East meets the West!
PRESENTATION POLICIES
SVP Abstracts are reviewed by the Program Committee and occasionally by outside
reviewers. Authors are responsible for the technical content of their articles.
Unless specified otherwise, coverage of abstracts presented orally at the Annual
Meeting is strictly prohibited until the start time of the presentation, and coverage of
poster presentations is prohibited until the relevant poster session opens for viewing.
As defined here, “coverage” includes all types of electronic and print media; this
includes blogging, tweeting, advanced online publication and other intent to
communicate or disseminate results or discussion presented at the SVP Annual
Meeting.
Still photography, video and/or audio taping or any other electronic recording at the
SVP Annual Meeting is strictly prohibited. (The SVP reserves the right to engage
professional photographers or audo/videotape professionals to archive sections of the
Meeting for the Society’s use.)
Editorial policies of Science and Nature magazine: If you are planning to submit, or
have submitted, your publication to Science or Nature, be sure you are familiar with
their embargo policies.
Please address any questions about program practices to the Program Committee or to the
Executive Committee.
Citing an Abstract in the 2014 SVP Program and Abstracts Book
This Program and Abstracts Book is an official supplement to the online version of the
Journal of Vertebrate Paleontology. The citation format for an abstract printed in this book
is: Journal of Vertebrate Paleontology, Program and Abstracts, 2014, <insert page number
here>.
5
November 2014—PROGRAM AND ABSTRACTS
2014
SVP Schedule of E
vents
(subject to change
)
All e
vents are held at the Estrel Be
rlin unless otherwise noted with an **
n t/ Fu nct ion TU E , Novemb er 4 WE D, Novemb er 5 THU R , No ve mber 6 FRI, Nove mber 7 S A T, Novemb er 8 on Des k 2: 00pm – 7: 00 pm ROTUND A 7: 00 am – 5: 00 pm ROTUND A 7: 00 am – 5: 00 pm ROTUND A 7: 30 am – 4: 00 pm ROTUND A 7: 30 am – 4: 00pm ROTUND A os iu m 8: 00 am – 12: 15 pm Archaeopteryx – T he Icon ic Fossil in Modern View HALL A 1: 45 pm – 4: 15 pm Th e In fl uen ce o f R. McNeill A lexan der on Paleobiologic al Inferences HALL A 8: 00 am – 12: 15 pm Putt in g Fossils i n Trees: New Metho d s fo r Co mb ining Mo rp ho lo gy, T ime , a nd Molecules t o Est im at e Ph ylogen et ic Posit ion and Divergence Time s of Living an d Fos sil Taxa HALL A 1: 45 pm – 4: 15 pm Eco m et ri cs : Q ua nti ta ti ve T ra it-Ba sed Appr oa ch es to Biotic Cha nge HALL A 8: 00 am – 12: 15 pm Th e L ives of Te mn os pond yls: In vest igat ions int o th eir Biology HALL C r Pr iz e Se ss ion 8: 00 am – 12: 15 pm HALL A arat ors’ S ym p osi u m 1: 45 pm – 4: 15 pm HALL D Sessi on s 8: 00 am – 12: 15 pm T echni ca l Se ss io n I Ce tr ti od ac ty la a nd Ma ri ne Ma mma ls HALL C 8: 00 am – 12: 15 pm T echni ca l Se ss io n VI Euarch ont oglires HALL C 8: 00 am – 12: 15 pm T echni ca l Se ss io n X Neogen e Mam m alia HALL D 8: 00 am – 12: 15 pm Tech ni cal Session XIV Mesozoic Rept iles HALL A 8: 00 am – 12: 15 pm T echni ca l Se ss io n II Mesozoic Vert ebrat es, prim arily Ma mma li HALL D 8: 00 am – 12: 15 pm T echni ca l Se ss io n VII Ar cho sa ur s HALL D 1: 45 pm – 4: 15 pm Tech ni cal Session XI Mam m alian Dietary Ecology HALL A 8: 00 am – 12: 15 pm Tech nical Session XV Gen eral Met hod s an d Ma cr oe vo lu ti on ar y Pa tte rn s HALL C 1: 45 pm – 4: 15 pm T echni ca l Se ss io n III Th eropod Din osaurs HALL A 1: 45 pm – 4: 15 pm T echni ca l Se ss io n VIII Paleogen e Mamm als HALL C 1: 45 pm – 4: 15 pm Tech ni cal Session XII A m ni ot es an d Ea rly Rept iles HALL D 8: 00 am – 12: 15 pm Tech ni cal Session XVI Fis hes an d E arly Tet rapod s HALL D 1: 45 pm – 4: 15 pm T echni ca l Se ss io n IV Cen ozoic Mammals HALL C 1: 45 pm – 4: 15 pm T echni ca l Se ss io n IX Sq uama tes HALL D 1: 45 pm – 4: 15 pm T echni ca l Se ss io n X III Paleozoic Fish es PARI S 1: 45 pm – 4: 15 pm Tech ni cal Session XVII Pe rm o-Trias si c T et ropods HALL C 1: 45 pm – 4: 15 pm T echni ca l Se ss io n V A m ph ibian s HALL D
6
© 2014 by the Society of Vertebrate Paleontology
n t/ Fu nct ion TU E , Novemb er 4 WE D, Novemb er 5 THU R , No ve mber 6 FRI, Nove mber 7 S A T, Novemb er 8 r Se ss io ns p : 7: 30 am – 9: 30 am Po st er Se ss ion I: 9: 30 am – 6: 15 pm Recept ion : 4: 15 pm – 6: 15 pm ROOMS 2 an d 3 Po st er Se ss ion II: 9: 30 am – 6: 15 pm Recept ion : 4: 15 pm – 6: 15 pm ROOMS 2 an d 3 Poster Se ssion III: 9:30 am – 6:15 pm Recept ion : 4: 15 pm – 6: 15 pm ROOMS 2 an d 3 Po st er Se ss ion IV: 9: 30 am – 6: 15 pm Recept ion : 4: 15 pm – 6: 15 pm ROOMS 2 an d 3 it V ie w ing 9: 30 am – 6: 15 pm ROOMS 2 an d 3 9: 30 am – 6: 15 pm ROOMS 2 an d 3 9: 30 am – 6: 15 pm ROOMS 2 an d 3 9: 30 am – 6: 15 pm ROOMS 2 an d 3 ine ss M ee ting a n d um 12: 30pm – 1: 30pm HALL A arat ors’ Me et in g 2: 00pm – 3: 30pm ROOM 4 l Eve n ts 7: 30pm – 8: 30pm Special Presen ta tion by Professor Hans W. Fricke **MUSE U M F U R NATURKUND E , LECT U R E H A L L 7 7: 30pm – 10: 30pm Welcom e Recept io n **MUSE U M F U R NATURKUND E 7: 30pm – 11: 30pm Stude nt Roun dta ble Forum a nd Reprint Exc hang e HALL C 6: 30pm – 12: 00 am Auc ti on HALL C 7: 00pm – 10: 00pm Awa rds Ce re mo ny HALL A 10: 00pm – 1: 00 am A ft er Hours P art y HALL C /D
*Ple
as
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is
no
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30
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7
November 2014—PROGRAM AND ABSTRACTS
2014 SVP Workshop Offerings
For Pre
-re
giste
red
Atten
dees
DayMON, Nove mbe r 3 9: 00 am – 5: 00pm Phylogeny, Evol ution, and Bioch ron ology of t he A rvicolidae Museum für Naturkunde, Pa le onto logi ca l Se mina r Room TUE, Novem b er 4 8: 30 am – 5: 30pm Paleopat hology – Est ablish in g Criteria for Recogn izin g Bo ne Pa th ologies Museum für Naturkunde, Pa le onto logi ca l Se mina r Room 10: 00 am – 4: 00pm T ip-Da ti ng: E sti ma ti ng Dat ed Ph ylogen ie s Usin g Fossils a s Term inal Taxa The Le ib niz He adqua rte rs 1: 00pm – 4: 00pm In fe rrin g Diet and Dent al Fun ct ion fr om Den tal Microwear Text ures Museum für Naturkunde, Le cture Ha ll 8
2014
SVP Field Trip O
fferings
For Pre
-re
giste
red
Atten
dees
Day THU R , Oct ob er 30 – TUE, Novem b er 4Famous V
erteb
ra
te
Fo
ssillagerstätte
n in
So
uther
and
Central Germany
Time: Field Trip will depart
from th e Frankfurt A irport at 12:00 n oon on Oct ober 30, an d arrive at 6:00pm o n Novem be r 4 at th e Est rel Berlin (h eadq uart ers h ot el) P ic k up Lo ca ti on: Frankf urt Airport Drop off Locati on : Est rel Berlin S U N , Novemb er 2 – TUE, Novem b er 4 Tet rapod s an d Fi sh es f rom th e Pe nn sy lvan ian an d Pe rm ian of Saxon ia an d T hu rin gi a Time
: Field Trip will depart
at 8:0 0 am an d arrive at 6:00pm Pick u p /Dro p of f Loca ti on: Est rel Berlin MON, Nove mbe r 3 – TU E , Novemb er 4 Messel, Geiselt al an d Ro m e: Hig hligh ts from th e Cen ozoic Time
: Field Trip will depart
at 8:3 0 am on Nove m ber 3 an d arrive in t he early even ing on Nove m ber 4 P ic k up Lo ca ti on: H ote l Co ro na in do wn to wn Fr an kfur t a m Ma in Drop off Locati on : Est rel Berlin S U N , Novemb er 9 Field Trip t o Rüdersdorf an d Niederleh m e End s: 8: 30 am 4 :30pm Pick u p /Dro p of f Loca ti on: Mus eum fü r Naturkun d e
PROG
RA
M
AT A G
L
AN
CE
Ha ll A Ha ll C Ha ll D Ha ll A Ha ll C Ha ll D S ymp os iu m 1: A rchaeopteryx – T h e Iconic Foss il in Mode rn Vie w Technical Sessi on I Technical Sessi on II Rome r Pr iz e Se ss ion Technical Sessi on VI Technical Sessi on VII WED WED WED THU R THU R THU R 8: 00 am O’Con no r Mih lbach ler Niedzw iedzki Yi Sch m erge Peecook 8: 15 am Dyke Roes sn er Pr as ad Viet ti Kraat z E zcu rra 8: 30 am Ra uhu t H ecke be rg Me ye r Schmi tt T omi ya Sto cke r 8: 45 am Kundrat Maier Marzol a Rabi Flynn Sobral 9: 00 am Erickson Bibi Jacobs Pim ient o Sm iley Dzikiewicz 9: 15 am Balan off Fait h Bell Kolb Bert ran d Foffa 9: 30 am Lon grich Cost eu r Mart in Jon es Morse Müller 9: 45 am Habib Boisserie Crom pt on Grass Silcox Gign ac 10: 00 am C O FF EE 10: 15 am C arn ey L ih oreau Hof fm ann Du nn B eard Man ni on 10: 30 am Sch w arz-Win gs Gin gerich Zh an g Cuff Kem p Wilberg 10: 45 am Lefevre Bebej Bolort set seg Clarke A sh er Baier 11: 00 am Mo yer Lo ch Co rf e But to n Co te Cer io 11: 15 am B hullar C orrie B am forth B urch C as an ovas Vilar Vin th er 11: 30 am Ben to n Park William son Brown A lm écija Padian 11: 45 am Mit ch ell Buc hh olt z Grossn ickle Brockleh urst Scot t U nwin 12: 00 p m Sm ith Crerar HallidayBales Wroe Dean
12: 15 p m BRE AK 1: 30 p m Ha ll A Ha ll C Ha ll D Ha ll A Ha ll C Ha ll D Technical Sessio n III Technical Sessi on IV Technical Sessi on V S ymp os iu m 2: Th e Influence of R. McNeill A lexan d er on Pa le obio logi ca l Inferences Technical Sessi on VIII Technical Sessi on IX 1: 45 p m Lan ger Macph ee Gao Palm er Th eodor Geart y 2: 00 p m St iegler Sán ch ez-Villagra Van buren Ch rist ian Secord Polcyn 2: 15 p m Fot h Fin arelli Ch en Rayfield Rose Cam pbell 2: 30 p m Ibrah im Weisbecker Baez Sn ively Fran zen Sim oes 2: 45 p m Gold Morris Vasilyan Preusc ho ft Gh eerbran t Con rad 3: 00 p m Hen d ricks Couzen s Zh an g Jan is San d ers Caldwell 3: 15 p m Laut ers Cook St ein Hut ch inson Sm ith Hsian g 3: 30 p m Fun sto n Beck Ma ddin O ter o Bor ths Wer ning 3: 45 p m Tan aka A tt ard Kawan o Sellers Haberset zer Mah low 4: 00 p m Varricch io Sh arp Boeh m er Falkin gh am Em erso n Kem p 4: 15 p m Poster Session I Poster Session II 6: 15 p m
8
9
November 2014—PROGRAM AND ABSTRACTS
Ha ll A Ha ll D Ha ll C Ha ll A Ha ll C Ha ll D Sym p osi u m 3 : P u tt ing Foss ils in Trees Technical Sessi on X S ymp os iu m 4: Th e L iv es of Te mnos pon d yls: Inve stiga tion s i n to the ir Biology Technical Sessi on XIV Technical Sessi on XV Technical Sessi on XVI FRI FRI FRI SAT SAT SAT 8: 00 am Mat zke Hooker Marjan ovi Morh ard t A lroy C oat es 8: 15 am Irm is San salon e Ru ta B ou rke Fras er Rich ard s 8: 30 am Wagn er Th om pso n Marsican o Maidm ent Madden Fisch er 8: 45 am Guillerm e Su Beigh tol Hill Mych ajliw Criswell 9: 00 am Po l Ri nco n Se ngup ta Ca mpi one Va n De r G ee r Un de rw oo d 9: 15 am Clarke Rabin ovich San ch ez Rom ilio Kilbourn e Sferco 9: 30 am Putt ick Sam on d s Kon iet zko-Meier Barret t Macleod Davesn e 9: 45 am Warn ock Lewis Mukh erjee Priet o-Marq uez Wat an abe Close 10: 00 am C O FF EE 10: 15 am Ksepka Saila Dan to Kay Clavel Clout ier 10: 30 am O’Reilly Goswam i Fort un y Zh en g Mabee Qu 10: 45 am Friedm an Tsen g Reisz Nair Pian Lem berg 11: 00 am Turn er Balisi St eyer U pc hurch Biller Clem ent 11: 15 am Wr ig ht Sha w Wi tz ma nn Fr on im os Ca sta nhi nh a Ri chte r 11: 30 am Broch u Wan g Pé rez Ben Moon Koyabu C lack 11: 45 am Gorscak Miller Miln er Maxwell Ch att erjee Mon d éjar-Fern án d ez 12: 00 p m L loyd B arn os ky Sch eyer Sch u m ach er Ferreir a C ard os o Po rro 12: 15 p m BRE AK 1: 30 p m Ha ll A Ha ll D Pa ris Ha ll A Ha ll C Ha ll D Technical Sessi on XI Technical Sessi on XII Technical Sessio n XIII Sym p osia 5 : Eco m et rics Technical Sessi on XVII P rep arat ors’ S ym p osi u m 1: 45 p m Clem ent z Turn er Keat in g Badgley Olroyd Beh lke 2: 00 p m Haveles Rich ards Scot t Bern or Sidor Brown 2: 15 p m Boch erens Leblan c Miyash it a Dam uth Sues Jabo 2: 30 p m Pat te rso n Sum ida Trin ajst ic St egn er Sh elt on Capobian co 2: 45 p m U ngar Verriére Rücklin Sch ni tz ler Hut te nl ocker Kn öt sc hk e 3: 00 p m Karm e Nich olson Sallan St en se th Fröbisch Van Beek 3: 15 p m DeSan tis Bever Giles Werdelin Jan se n Fit zgerald 3: 30 p m Rivals Lyson López-A rbarello Frit z Laaß Fox 3: 45 p m Reed Prit ch ard Ch evrin ais Meach en Walt he r Davidson 4: 00 p m Davis B uch wit z B razeau Head O’Meara E klu nd 4: 15 p m Poster Session III Poster Session IV 6: 15 p m
New edition of the world’s leading text
fourth edition
Completely revised and updated, the fourth edition of Vertebrate Palaeontology takes into account the developments over the last ten years to present the latest research in the field.
Designed for palaeontology courses in biology and geology departments, this leading text will also be of interest to enthusiasts who want to experience the flavour of how the research is done. The book is strongly phylogenetic, and this makes it a source of current data on vertebrate evolution.
Study aids include key questions, research to be done and recommendations of further reading and websites.
Read sample content and order at www.wiley.com/go/benton-svp Save 20%! To benefit from this exclusive discount make sure you quote VBJ15 at the checkout. The discount is valid until 31st December 2014.
Exclusive
Delegate Discount:
Save 20%
Vertebrate Palaeontology, 4th Edition Michael J. Benton
Professor of Vertebrate Palaeontology at the University of Bristol, UK
October 2014 | 480 Pages | Paperback ISBN: 978-1-118-40684-7