Gomes
3e Lydia Huguenin Queiroz
11Departamento de Solos, Universidade Federal Rural do Rio de Janeiro, BR 465, km 7, 23890-000, Seropédica, Rio de Janeiro, Brasil. 2Departamento de Fitotecnia, Universidade Federal Rural do Rio de Janeiro, Seropédica, Rio de Janeiro, Brasil.
3
Programa de Pós-graduação em Produção Vegetal, Universidade Estadual do Norte Fluminense “Darcy Ribeiro”, Campos dos Goytacazes, Rio de Janeiro, Brasil. *Autor para correspondência. E-mail: arcangeloloss@yahoo.com.br
! "# $% # & ' (
' ) * +,( - ! $. *- # * # "# #
# # # '/ ) # # / # # / # #( # '
0 ( # # # ! # ) # & #
! ) # +, # $% # ! ! # # 1#
# . # *- # ) $% # 2 # # #
# / # # # / &
# - ) $. $. +, - 3
+, $. # # / ) # / # #
) # / # # #
-! " # $. #
$% & $'& () * ) * ' + "
( **# # # * ( ,) $ ( - %$. 4/ # / !! !
/ # ! & ' 5 ( ' ) +, (
/ ! ! # # / # ! # # 6 # '/ ) # 7
# 7 ( ! 7 & 7 ' 0 ( 4/ # 6 # 7 )
-7 / & / 7 / / ! , 7 /
+, 4/ ## ## # 7 # ! /
)# ! # # # #/ # / # 6 # / 7 # #/ 7 / ) #
/ / & # # #/
## ! +, + / 7 / +,
/ ) # 7 # 6 # #/ 7 / ) # # # !
/#0 + ( 7 /
# # ##
8 9 # : # # ! *! #
! # # # #
# - # $. # #
# ';< = 5 3 0>24? @(
$. # # #
! # ): /
9 $. 9 - #
# # # # ) #
# ## # A # #
$. ## # # ' 3B2>+ 324?5
3B2>+ 5 2CC+ 3+ (
0 ' ( A
) # & / # 9
$ D A&
3 A # # #. #
) # ! # # ! / # #. #
# # # #
) # # / # E ! *!
# # ! # 9 # / #
1# # # # # ! / # 9
# ) $. F# ! #
/ ) # ' ( ! $. #
# A A # )
. = # # ! $.
9 # # . #
: ? # A
$. # ) # #
G # # # )
'>?C23H+5 ?IH (
? # # # 9 #
A # $. # # - #
# # # # #
! /
9 9 # ! # #
# A
$. # # ) # # !
$. # # ##. *- # '= +J 5
B? 2 (
G # A # $.
# 9 A A )
8 # ) #
# # $.
1 # # # *
A ## # 9 .
# ) ) 9
& '= +J 5 B? 2 5
, C,? ( # # # #1
# #
## A 9 - # #
!A # 9 - # #
', C,? (
= ) # #
9 9 K - )
$. # 9 # . !
! !
'L 3+0; ( # 9 # ! #
## ) ! #
# M # - #
# - A
$. * ! # : 9 #
$. * : 'N C4 33
(
# # # # ## ! # 9 F
# $. # # ) # #
9 # # / # #
/ ) # # / # # # / ) #
' ?IH ( 2# # / # # # 9 #
9 ## # ! # #
# # # 9 / ) # # 9 " #
# # # # # / # #
# / ) # # #
# & # ' ?IH 5 +3 ?CH 4? O (
4 # # # & # # A #
# ##. *- # $.
# # $
M # *- # A
! - ',?>+ 3+5
= +? @( ? ! A 9 #
# / # # . #
/ # # ) " /
9 # #
! 'C2+ (
? ) * ' +,( &
# A A # # # # M 9 &
1 A ' + ( #
# # / # # #
- ' +C 3G D5 2CC+ 3+
5 >? @(
# # # " # - # # #
# #
! # # A # # # # F#
! # $% # & #
# # #P B - ' (
-= ' O( ) # ) / # #
# # ' # (5 C / ' D (
) / # # #
/ ' (5 ; 6 ' (
C ) ' ( # #
9 # ' (5 C /
' D)( ) / # # / ) #
/ # # # ' (5 3 #
' ( ) / # # / ) #
# / # # # ' (
3 . #. # ) / #
9 # ## ! ##
- # # # ##
# ) / ): #
! $. - ) $. # #
0 ! #
$% # ) *
! !! !
? & ! - = 9 #
L # I # C C
L 'I CCL( 1& + #
'+ ( # Q DOR
D Q D RS B 7 #
* 7 ## ! $.
;T
?# # # ! #
# - ) D - #
& 9 # #
#
G ## ! # # #
# @ # # "#
#P # # / # # 'U V (
# / # # 'U @ V (
/ ) # 'U @ O V ( # ! / #
! # # #
* 5 # # '
) # ( # "# # #
* "# #
# - # # #
1# ## # # ! # #
& #
# 9 # # 9 ! - #.
# # # # $. ! ') (
# +, > #
# ## $.
) * ' +,( O W ) $. #
$% # +, = # # #
! - # ! & # - # # 9 !
# ) # # $. #
#.
# # # ! #
- F # ) # = 9 #
L # I CCL . D I - #
# )# # !
# 9 # F ! X # /
# # # ) # / #
- # # / # '
# $ ! O # # / #
# & ( 0 '
> +,( # # #
' "# / # - # #( # /
$.
A ! Y0 . !
## # $. # # # / #
# * # $% # ! ! #
# 1# # 9 ! #
. # *- # ) $% #
= ## # # # ! # #
. # #
# = # !
# $.
! M # # 9
# # # #
? & - !
) # # "# $% #
! # * - # # #
& ! #9 ! & ' "# #
# "# $% # +,(
3 # # ) # !
# # '> ! #( / #
V # '0 / , ( # ##
#! $. # # ?# # # !
# ) # F # V $.
4 # # # A # # #
# ; OW ) )
" "" "
?# ! # # # # $% #
) * ' +,( # ! $ #
# ! # ! $. - ) $.
'4 ) ( 2 $. F ! $. #
# / ' $. ( # # #
# # # # / ) # / # #
# # # / # # # # # F
$. > # # # / ) #
# / # # #
$. +, 9 #
# # / # # ! # # #
$. > +, 4 )A #
)# 9 $.
+, - # # # / # # #
9 F#
#
& ,# 2! $. # ! # # #
$% # Z + ) * ' +,( $.
! $. - ) $.
0 - I CCL A 2# C
L . D
4 # # #
0 $.
N ) / # > / #
J #
# #
> W
O D ) OO
@ , O D ) D @ 0
0
@ 0 @@ ,) @ ,
@ , D 0
D ) )
C
-0 D ,) D ,
D O ) D O ,) ,
O , DD D 0) @ D ,
, $.
D D ) @
A # # # # M# . ! # !
# $% # # # ; OW5 # #
# M# / . ! # # # #
$. ; OW
' ( # $.
# 9 - A )1
' N??; ! ( # #
# / # # ! # $% # +,
' D ( . )#
- # # # # $% #
$. ! # # # 2#
A ! )# #
) / # # )# $% # # ! #
# "# # # # #
0 # $% #
= - # "# # # #
# / # /
$. 5 # # A # #
# # # / # # ! # # # F
# / # # # / # # #
- #
$% # +,
= ' ( !
) * ' O > (
-OW 'F ) # ) " ) OW
! A ( - # #
/ ) # / # # ) - #
9 # # # / # # # # / #
# # $. F# # # / ) #
# ) " 2## A # /
# #
2 $. F ) $. # # # /
# / ! ! $.
# # # # # / ) #
# # # # # F
/ # # )A )
) $. # #
= #9 ' ( A ## 9
/ : ) $ 1 9 #
& # ) # # # :
9 *) # ) # ##
$.
G ## ! $. # ! #
# F# # # ' $. 0X3
! # 1 # ( ## ! #
- ' ( #
F ! $. *- # * # N
/ 1 # 9 # # A # 9 ## !
! *- # * # # #
## # ! # # # :M
) # # # :
# $. # # )# V # 9
- F ! $. *- # 'N C4 33
(
= # A 0
# ) $ 1 9
'= [I > ( $. # #
! # # 'N C4 33
( : 9 # # # W
- # # / # # 2# # # #
#. ) # ' ( #
$. # 9 - A # #
# / # # ! # $% # +, ?#
# 1# # . )#
-# -# #
? ) * #
- $. > # #
/ # # 0
! - /
) $. # . 2# #
/ # # # # ! # 9 /
#
0 # & #
- ) $.
$. +, - 2 $. F
) $. ! )# 9 # # # # / # #
/ # # #
$. +, >
" # "" ## " #
3 0>24? 5 32BC2>>2 C C , 5 ;?2N>2C N
B $. '> ( B # )
', ( ! # # )# # # 1
& & $ # )5 $ 50 @
3B2>+ 324? B 5 3B2>+ , > G = #
#P # # # $% # #
% # (# 6 ) ) 5# O
, C,? L B ' 7 #5 P $. # K
! K # K
/ 8 J $ # P
,?>+ 3+ 0 5 = +? J C 2! #
# ) # # ) *
- # # # '
> ( ') ) $ 85 O
@
2CC+ 3+ = 5 ,?C4?>+3+ 5
HI 2>> 4? C+, ; 0 5 ;?2N>2C N
= $. # 9 - A )1
' N??; ! ( #5
' 9 $ : O D
BC H+ 3? 4 4 5 =+J244 ; > 5 =2C2+C
5 , 3H 44? G 2! !
$% # ) *
-# # ! / # # ' # ( \C ##\
' # ;* D
N C4 33 N 45 ;2 42C G 2 5 G J+ L]3+?C
4 5 B232J2 C > < < P #
# 3 7 L # P = N
N C4 33 N 45 ;2 42C G 2 5 G J+ L]3+?C 4 5 B232J2 C > < < < < #
( < # 3 7 ^ 6P 2 7 0 #
L 3+0; L $ 9 ( " ) ) C L P , # #
;< = 3 ()23 5# (# <
5# B *) P
;3 =+; L B 5 HI 2>> 4? C+, ; 0 5 0 C= 32HH+ 5 4 J C2 C 5 ;?2N>2C N
+ ! " E / $.
) * $. # 9
'0/ ( 0 '9 # ( "# # #
% O@
>?C23H+ N 5 ?IH N 5#
% ,) = "# ,: # # #< (# 3 ? ## P =
>? 5 42+_2+C , 5 I3`a? B 5
N + = B 5 >?IC2+C? G 0 5 ?IH L C
2 - # # > #
Z + , * ' +,( ' 9 $ :
D @
+C 3G 0 5 0N > I3 3 3 L 5
N? 33 5 GI4C > 5 0?2>N? B J
2 & * +, ! # #
- # # / # # # & #
## b 6 7 R - ' 9 #
$ # @ D @ @D D
32J2 4 5 0 C= 32HH+ 5 HI 2>> 4?
C+, ; 0 5 C230? C 2
# ! $. # $% #
# - # # >) $ <# ) % # D O
= [I > 5 0N > I3 3 3 L 5 C ? L G 5 J >2 C 5 +>J 0 C @ ) ) ) 5# < < 23 (# < * ) ;*# > #P I > X = I> > 5 ,?>+ 3+ 0 5 0?CCc > 5
02>?4? + , 2! ) *
-- # # / ) # / # #
) - $ # )5 $ 50
D D D
C2+ L C 5 0N > I3 3 3 L 5 >+ > 0 ? 5
>+ > 0 2! # ) *
- # # & !
G ' 9 $ A D D @
C+,2+C? 3 ? 5 = +J = G ? 5 +>J L 0 , 5
= +J C 2! ) * # ) # #
# # 9 # '
0 ( # % # (# 6 ) )
5# @
C?0N 0 5 [I2+C?H L > 5 [I+ 23 C 0 5 HI 2>> 4? C+, ; 0 5 >4N I ?44 33
2 - # # /
' '= / ( G G ( ! #
$% # ) * # $.
') ) $ 85 @ D
C?0N 0 5 [I2+C?H L > 5 HI 2>> 4?
C+, ; 0 = $. #
A # 9 # $ O
D)
CH+ + 5 =+J244 ; > 2! # # $% #
) * - # #
# # ' # ( ' # ;*
@ O
?IH N ?# # # # )< #5#
$ ; D
?IH N 5 +3 ?CH 4? C 2
-# # " > ) =
/ 8 # # % @ O @ O
# $ %$ &''( %&$ &''(
> # ! P 4/ # # ## # ) / # ! /
0 0 # ) > # 7/ / # # # # )