Morfologia e distribuição de Paralia Heiberg (Coscinodiscophyceae) no Sul do Brasil.

#$%&'('!)'*+,-,./,1'! 

0123415

1678

9:;8

;<

=>

2<

?@>

<

1:8

1A

8

BCDCE

F

C8

GH<

?H268

I

JKLMF

NKOF

LMKPQRMSCST

8

<

:8

=1@>

4H2:8

U29V<

5

W

092<:X=8Y92Z<9[\8]^_<98`9569>><a\8b1=H5<8c1@V9d01=<e9::f8g8b1=H59:H8h9@;92H=dc<5_9f

_{&&'((&ij (&()&)7}

& &k(5&)&ij  ''&l&7m'('&n&9&&o&&'&

_{m'('&n&&(7o& )(m'('p'o&jqj7}

r&&  s't&7(uv7o2w""w""7&(7397 &(' &'(x)*)!

_{& &k(5&)&ij  s't&7m'('&n&3'5&9)}

6 5i&(w""7o2wy"w""767397 &('&'&&&'x &'t

y_{m'('&n&9&&o&&'&7o& )(m'('p'z'&7}

r&&  s't&7o2{{"y"w""7n'&k'(79o7 &(' 3t!' "|00"6t' "|00"

6 9z36oz}z$ $%&'('!)'*+,-,./,~€‚-,‚,9&ƒ&'„4&) &+…†‡.ˆ,‚, ‰2$!Šo†ƒ()'ƒ'$'&(*"(& (* &‹&(&%(' 7ƒ$'t$ ƒtt&$t&(*9&&o&&'&&3'5&9)(&(3&' $%7 $ &''&(t' ('*** $4$1 '($(t' (7') !*t('$ t$&'&') !*'*($&*&t*(&&'&( $(t'(ƒ '$'$&'&!)&t&$%$&!*)'' (ƒ'$ƒ'ƒ& &) ‰vo{voŠ&(&''%‰w"Š&(

Œ%ƒ('& (7+,-,./,†~€‚-,‚,Ž†+,-,./,†‡.ˆ,‚,

329m#}012A1516<98H8;<=>2<?@<‘18;H8BCDCEFC8GH<?H268IJKLMFNKOFLMKPQRMSCST8:18c@58;18U29=<5’8 6 *'&'('!)'ij+,-,./,~€‚-,‚,9&ƒ&'„4&) +…†‡.ˆ,‚,†‰2$!Šo† *& ()&&( "& (&(st(' (&((7(“)&'(*& t&(& &t(&((&(9&&o&&'&3'5&9)o &ij” *'&7((t' ( '(&( &(''* (t&(1 '(*'47't'&  •  t)&(&t&'&• &'(((!&*&t&&&(&&((&&ij6 !&(&( (t'(*& &&( &&&!)st'&&'&*& t&&( & !'(t & & &'&ij &)&‰vo{voŠ(&''&‰w"Š

&&&(t$&'& pt&(7+,-,./,†~€‚-,‚,Ž†+,-,./,†‡.ˆ,‚,

0

4z3rmoz0

4

z$)(+,-,./,1'!$&($' &– 't&%&tt(t'(†6 †$(7† +,-,./,†‡.ˆ,‚,†‰2$!Šo'(&ƒ'%‹ƒ &t'(t'(0$&(%&(75&t'&‰""Š &&ƒ(t'($)(7+…†../—‚/ˆ,5&t'&7 &!$'t&–ƒ'$''t&)'*)'$ &'(&%!&t$(*$9&*9&&o&&'& ‰v9&˜9Š6*ƒ&(79&ƒ&'‚†,.…‰""Š (t'!ƒ(t'(*+,-,./,+…†~€‚-,‚,9&ƒ&' „4&) &+…†ˆ,—/‚,‚,9&ƒ&'„4&) 7*) '™&&(ƒ&(#t%7#&t5''&%„ Œ&t &‹&‰"Š(t'!$(t'(ƒ'$' $+…†.š€›/—/€,Œ„™&(t'(t – ‰+…†,../š€//#&t5''&%7+…†ˆ-,œ~š-//#&t5''&%7

&+…†ž-Ÿ,€//#&t5''&%Š*)'™& &'t&7 && &7#–'t7&1&ƒ&''

+,-,./,†~€‚-,‚,ƒ&(*)'&‹& )*™&&((& (*&''!&%‰6'&‹ &%7v4Š* &!&t‹'($'&&‹‰Œ($'‹'0(&7

v4Š7ƒ$'+…†ˆ,—/‚,‚,†ƒ&(((ƒ'%'('!)

&t&*‰0($'&‹'0(&7yv4Š0$(&  '79&ƒ&'‚†,.‰""Š&(*)+…†‡.ˆ,‚,!) $$(t'(*+,-,./,'tt)'$(&  t&''(&(& (z$($(t'(*+,-,./, '**'(&&'('92#7&$' (' &&&&('%&(((('q#

+,-,./,†‡.ˆ,‚, &%(&' & &‹t '(&&&(&)$ ƒ  (ƒ'(!%1!(„ #t )'‰wwŠ73*(‰w{yŠ''&

 563#067$7%6&56%%07'79()*6$(90$6447_{3+&6)'632990&,673}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

QRSTUVWX'/1/!D/Y;BLB@ZB=IJKJ>CLBCB;<=>?@ABCBM -1Y9..#../..3/5.9D1.1/1DM ./.F.1/ 1. 1 ./.!D.-Y8[.M/M1.//0 /9DM //1M#D !/.7..D !1 YM/11-/1M.-0!YD/1. !05!\MJC=B@4""H: 9. .//..DM1M./1 ;<=>?@ABCB/M/1. 17!DY]M./ /D1./1P/./.!/D1./14/ .]/1.&$ /-.M1:.YD/31. 4H6:7 D1.%//4H6:7P^-/P.1 JC=B@4HH8:79D.$1/ .JC=B@4HH:79D. $1/ .+311(//.4HH6:7$D.JC=B@ 4"":7$./-JC=B@<4""6:7.$./-JC=B@< 4""H:2/ ...M.!/1 YM1.1Y. !D-4$D.JC=B@7"": .3/5.9D431.7H6: ;BLB@ZB=IJKJ>CLBCB].1/D1.]/M&$. 92$!1./1!09D.$1/ .JC=B@4": Y &...#-/2N7!D/ . ..].1 %M./ YM/11D0].11 M /-.!1./1;<=IJKJ>CLBCB=.=;<=>?@ABCB= !.1&$.92$Y .//.1-/ 1 - ./M ]/M1-/ 1/D1./M /.D7./1/.M//1/!D/. ./.!D.-/M].1Y1DM ./

$6%230

6&

64'

$2%1('9

9. /]-./D/9..#../. 01.4P/._/179..#../.9.49#:7 5"79.5"7986579.86579:%M 1/1]1.!/1M`O.9D73.1 5.7.5D.313.15..5D.31. -./.1 /-1!.04 .^.4:.M0 abcdefgcbdhicdjfkecflcdmndhomdbjpcbqdafgdrafsbmpcqd 43.1.0.-D!//:7MMM.7 O.9D1//1-./1M%MMY M1. /-./5D.31.!]. 8 7/M3.15.M././].1!] 6." 7./O.9D!]. 8 0.7.F1. 1]-- M0Y G.D.0""G.D.0""67. tucgdojhidnmbravgciwgcdahdxyz

0.//M1. /1-/!.!7 81!./D 1/1Y 1. 1--M -.11Y9..#../..3/5.9D1.1 M.1"0.1]!1D1. M/1/!D/.Y{D-0Y;<=IJKJ>CLBCB=. ;<=>?@ABCB/1DM ./4%.!:6M1 !.-M1.|D..01/ 1..1D!}-  /-/17 ./01 //D.]/M. . .Y.D"6 .. .~/ D /Y  4%JC=B@<"""€/+$17"":

#$%&'('!)'*+,-,./,1'! 

09&&0&&'&7(& (122)(' 34567489:4;<4<6=>43?6@7A@6BC4DE6F4GH7>4IJ@<3KL $%&MN&O'$*'&60&((' !&2$P3'5&9)39Q7&O(& ( 1221'$&2'2&P"R  ($ &)Q7(1'$MN!)*** & F6>SF6435F4T6?74H54UK3==4V3=T=489WW4<XBY4Z6FH<6574 (& (* $2&(*9&&0&&'&9&1 &O* $(&($!%(2&'$() D[H3K4=35F4A57HK494<<4F6?7>Y4Z6FH<6574G3=4T6?74 H54V3=T=4GH7>49W4<K4JI4\]4KAUJK4=JKA7HJ5435F4 &MN* &$%1&(&9& '('(

1$$(2'(1!(&(' ^')

_$(& (* 9&&0&&'&9&& '('$1!&') *$`'('& ^&9&&0&&'&P^ab3Q71!&')  *c6&'2$3192$)*$#)() 0'd2'&(4&)&'(&^)&efg!h'2& 3'5&9)P169Q&1!&') *$ `'('&^&&(P2aQ71$&($ (& (* 3'5&9)9&&(' '$1!&') *$`'('&^& 3'5&P1`35Q

ijklmn9& '('(1$+,-,./,opqrqst-,t,PQ+uosv.w,t,PQ1*)'9&&0&&'&P90Q&3'5&9)P39Q (&(7()$ &'

# 563$067%7&6'56&&07(79)*+6%)90%6447_{3,'6*(632990'-673} & ./.0.1123 .0/24 5501063.05.75/.707.8. 9/94 9..$..1.00.69$9.50.. .0531&.!&5 ./3.0 .0/ 315.!!0616$1":9.0.112 /01.;/;112;616$1""9 9/!0. 04 .;5013;./01 5;51</0;1!!291 06=:9. 45 .1.3.0 /..0001 3154.5.>?_{ .@3 .010} 0. 7.>. 1315.)2 /0 A"15 1;0; B9;.12;%1;0;2692%97 ;.0;1 0310/!0.;. 315.:" 64""0;0/015 .; 9$(""0/;.&50/!03!0 /01.C2)'="'- 1;0;..;;.1 .04"@-&53@110.;3.0" 

DEFGHIFJKFHGLFMFGNOPQRKSFTGJUOQRKQVGWXXGYPZYFHG 01101 .51.1.!/.; &5.1.!/.;..4.;50;104  3.05.75/.70.8.9/010 69..$..1.00.9.05315B1/

329*'&9

64(

(0

9$*990

)4

[\]^_\`\abc\dce_fceghg ijkjlmjnopqprskjsjt9.3.1,4./ 752;1.7 WWuvwXxGSVHyGwz{WwyG""

6B10"9 |}~}€}t‚ƒ‚„…~}…}t3.04/4 1;5.10 3155424/;06†#7..†= SZP‡FQRˆGUQG‰PQRPGŠPRPMKQPG‹HZPQTGŒ‰ŠˆxGLEKZF 210.4/0/034/.$.001!.;5 63996B1099..1.00 010.5.1. .@105.4.;. ;. 13.2!35. .1.; 4>5. .1510.0134 IUMFHGŒKVyGzˆyGDEFG‡PQRZFGEPHGLKTFGPMJEFHGSZZFTGLKRE WGRUGvGMULHGŒQURGMFVOZPMZŽGUMVPQKFTˆGUGSQFGIUMFHG 4.>1 .250. 016B19..14 1 /.5 .6B1#90;..2 .000./.0106B1=9./5 . .1.3 .@105 .4.;6B1"9&51. 450;1 0 4/9..$..1.00..1!3" .‘ 6†:9.0.105/.7073.0 5.7.8.9/735.0. .1.9/7 '...$;1’“731!1“.05.7.31!1“ B..713.0‘ 6†"90 10/.5.1.13.0"=” 6†:9 &5.1. 10.$.001!.;53.0 :” 6†9 •–a—c˜3.1.!/.;4|}~}€}t‚ƒ‚„…~}…}7|™t„š›}…}. |}~}€}0630;105355.1.!/.; 3.0005.œ9159..$..1.00.79$63593.0 5.91ž595/.7091ž998.9/919

#$%&'('!)'*+,-,./,1'! 0

12345678943:7;53<= 36>0?7964@6A6@630467 7A&5&70B""7C2?DEF>' !(70007 C0A40GDEF>' !(7000""7C2?0EFH'&I'(7 &'& 9)7B007CH?J3DD0EF &'&9)7B""7CH?J3 """"EF3'!'K&0$&70000DG7CH?J3D"DGEF ?&&&AL'MK7000""7CH?J30"00E=5)&N(7 0""7CH?J3""EF5)&N(700""7CH?J3""DEF5)&N(7 000""7CH?J3"EF5)&N(70O""7CH?J3"GEF5)&N(7 O""7CH?J3"EF5)&N(7O00""7CH?J3""EF5)&N(7 O000""70B""7CH?J3"EF5)&N(7B""7CH?J3 "0EF5)&N(7B0""7CH?J3"EF5)&N(7B00""7 CH?J3"EF5)&N(70""D7CH?J3"DEFP&9)7 0""7CH?J3""EFP&9)700""7CH?J3""0E P&9)7000""7CH?J3"EFP&9)70O""7 CH?J3"EFP&9)7P&9)7O000""7CH?J3 "GEFP&9)7O""7CH?J3"DEFP& 9)7O00""7CH?J3"EFP&9)70B""7CH?J3 "EFP&9)7B00""7CH?J3"EFP& 9)70""D7CH?J3EF3&(5&70""7CH?J3 ""GEF3&(5&700""7CH?J3""EF3&( 5&7000""7CH?J3""EF3&(5&70O""7 CH?J3"EF3&(5&7O""7CH?J3"EF3&( 5&7O00""7CH?J3"0EF3&(5&7O000""7 CH?J3"EF3&(5&70B""7CH?J3"EF3&( 5&7B""7CH?J3"DEF3&(5&7B0""7CH?J3 "EF3&(5&7B00""7CH?J3"GEF3&( 5&70""D7CH?J3"E30J5364Q2QJ9R?73' 5&7A&(('70O""7C1R35"0D6E

STUTVWTXYZV[T\T]C2$!EA7 '$&' ^9(_&O(_&(6_& '(1&'&7 CE=D`,../abc..,]de.f,g,2$!74&) *(L$H)) '7D

CH'(E

+,-,./,]de.f,g,]h&(*)* 'L$&'(h'$ $h%('iL(Cj7&&jL( klmnopqrlstunvsntwlnxyplnvzn{qstqn|qtqmxsqn}{|u~n h$'%'(&*)()(h*)A&((' !&L$CH'(E9&&'&(( ('(&$&'& &_'($&*&L& L& 'h&%!h$& &'&L CH'GE4i$& &'$&h(' h(*(0L&&%&($&('(&) $& &'&h &_'($ &*&LCH'E0L_'('(&($& '( &((&&'*' )&'( ($ &CH'E@$i& &%*$ &'$(&&'&'L&&% &(CH'E$&(&&'&!) !*(h'$& '('L($&&')&'('!)'@$'&  *$(L' (*)9&&A&&'&0(&C9AE &'!h&" CjE&(&'( 5)&N(73&(5&7&P&9)7h$&( & &'&9)7?&&&AL'MK73'!'K&0$&7 &3'!'KH&&7'h&( CjGEJ  '()&C9AE7$&'&'h&("€  CjE@$&'& '('3'5& 9)9&CA&(('!&L$Eh&(0€ CjGE

1234567893:7;53<‚9 36>0?7964@6A6@630467 76&MN7B""7C2?DEFA&5&70B""7C2? DEF>' !(70007C0A40GDEF>' !(7000""7C2? 0EFH'&I'(7 &'&9)7B007CH?J3DD0EF B""7CH?J3""""EF3'!'KH&&7O000"7 CH?J3GGEF5)&N(70""7CH?J3""EF5)&N(7 00""7CH?J3""DEF5)&N(7000""7CH?J3"EF5)&N(7 O""7CH?J3"EF5)&N(7O00""7CH?J3""EF5)&N(7 O000""70B""7CH?J3"EF5)&N(7B""7CH?J3 "0EF5)&N(7B0""7CH?J3"EF5)&N(7B00""7 CH?J3"EF5)&N(70""D7CH?J3"DEFP&9)7 0""7CH?J3""EFP&9)700""7CH?J3""0E P&9)70O""7CH?J3"EFP&9)7O""7 CH?J3"DEFP&9)70B""7CH?J3"EFP& 9)7B00""7CH?J3"EF3&(5&70""7CH?J3 ""GEF3&(5&700""7H?J3""EF3&(5&7 000""7CH?J3""EF3&(5&70O""7CH?J3 "EF3&(5&7O""7CH?J3"EF3&(5&7 O00""7CH?J3"0EF3&(5&7O000""7CH?J3 "EF3&(5&70B""7CH?J3"EF3&( 5&7B""7CH?J3"DCEF3&(5&7B0""7 CH?J3"EF3&(5&7B00""7CH?J3"GEF 3&(5&70""D7CH?J3"E30J5364Q2QJ 9R?73'5&7A&(('70O""7C1R35"0D6EF0B""7 C1R35"006EFB00""71R35C1R356E

ƒwlns„r…lmnvzn†lppynklmnopqrlstnxsn+‡]ˆcbcdg-,g, '( L %$'$$&'+‡]de.f,g, 9&h&'cg],.‡n}‰ŠŠ‹unmlkvmtlŒnywvmtnopqrlstyn}n "L(E*+‡]ˆcbcdg-,g,h$'h$&*) opqrlstynmqsŽxsŽnxŒlpnzmvrn‘ntvnn†lppy’n“vmn +‡]de.f,g,7A&h*C00ELL(C' q”lmqŽlunklmnopqrlstnqsŒnlnv…ylm”lŒn••n†lppyn}xsn q”lmqŽlunklmnopqrlst’nƒwlns„r…lmnvznmxsŽnvznkvmlynvsn $&*&L*$(&&'&(h&('** !h$(L'(C@&!E+,-,./,]de.f,g,$&(& )!'*('L&(h'$$('' !(!%9&h&'cg],.‡C""E&#L–)'— 4!C""E@$)$7'$H'!%A&h* C00E7$($h(&(L' $&$&((' '& && &($ &''&(L' (0$(&  h&%7(&&'&(*+‡]ˆcbcdg-,g,*  &' ($h&('h*($&*&L!) '$'&('(h'(h(L'!9  'L_'('(*+‡]de.f,g,]& ( $&')(&!%9&h&'cg],.‡C""E&A&h* C00E

0˜&&(h&(7+‡]de.f,g,(&( & ŒxqrltlmnmqsŽxsŽnzmvrn™ntvn‰‹n€r~nwxpln+‡]f,š/g,g, &+‡]ˆcbcdg-,g,&L$h'' ('(7&' zmvrn™ntvn›Šn€r’n{kl†xrlsynzmvrn{qstqn|qtqmxsqn 0(&$&h'&''& (=+‡]ˆcbcdg-,g,] &* " &&($&'&' * " h&(!('+‡]de.f,g, Q'&  &() (*]+,-,./,]ˆcbcdg-,g,h L('(h'$'(% &'&&+‡]de.f,g,] (&&](' '&$&L'!%9&h&'cg] ,.‡C""E

" 563#067$7%6&56%%07'79()*6$(90$6447_{3+&6)'632990&,673}

-./01234561&789: 7:;: 7:98;<=>?>@A>BCDEDFG?>G>H31577I;8I7 8 99 !J;9I;<99;99!J7!;K9ILHM1,97I<;7!7 9HN1OPQRSTOUVWXYZTO[\TQORZYO]Y^WSWT\RZO_WV_YOWZ`OabcO]\dV\ZeO_WV_Y]fOg129997979 7I;77<999h8 97K!9:i9IL9 9i7;89<I9J !I9:9 97KI879ILHj1'97;<h97 k99k;K9I;LH l199979I78k::7;7;97<999h8 976I ;8I;8h97 k99kHm197!79;I78;9k9;7;K9IL56197!7 9I78 9i7;9<9:KI879IL9nI;<;8 9K!9:i9IL 9:9!9;o-./01234NOpOqrOsXfO-./012gt2lt256OpOuOsXfO-./12jOpOqOsXfO-./12mOpOaOsXv

#$%&'('!)'*+,-,./,1'! 

0123456676849'$&: ':(: ':&$(*+,-,./,;<=.>,?,@664ABCDEAFGACAHICJKLEAML ''N@6O4ABCDEAFGACAHICJKLEAPQFRMLSATKDUAVIFPKIUACDDCLSKWADMXPAYCDDFRZ[A6\4ABCDEAFGACAHICJKLEAFGA &!):(@6]49&&'&'&'N('N'(*&&&^!&:_&N` & &_( 'N&%!N&:& &'^N$'&N`@6a49'!'&'&'N@ 6b4c&'*&$'':_'('^&N`@684AdKECMIAFGACAHICJKLEARMEQAVIFPKIUACDDCLSKWADMXPeAfVCIKA !&(g0123456676\AhAijAkJ[A0123456]l56bAhAmAkJ[A0123456al568AhAnAkJe

 563#067$7%6&56%%07'79()*6$(90$6447_{3+&6)'632990&,673}

'-../0!1!20/-023!  230-!49313-56789:"";33!30 3-42 33-3-0<=7>5?5@6A8687 2 3.033-30001 BCDECBFGHIJCKFLMNJFKCOCHGPFQRSTUFHVLKFQEVNFHCWXPGHPYF 3-;.0 30.3Z- 34203  0-0<=7@[9\8687033-300 3/2030.-0-/0-03-]34 -0-!]2 3:9313-567897""; )-2074<=7599^_6^\8`7<=7>5?5@6A8687 3<=7@[9\86871-0. 3-=7%20 20/-03-.. 3-4]0/3- /-/ -/0/4:0%3!;

abcdebfgdbhijklfgimlinoklimkoipbehiqoidlrkmldl 63400.2- 303- 0-33 -!4 3--3 3-30]023<=7 >5?5@6A868-01-4-0-!]-0]2 3- 3<=7@[9\868 34//]1-2<=7>5?5@6A868- 032330!0-0933#33-3003 :9#;:%3!;%2--1.<=7@[9\868.  2 3.%330-:stu93vts "uw;-]03!4xy/-x30+9]3 $0- 3:z{{RUFIGEF|CFMBCENMDCBFGKF<=7>5?5@6A868 %2-3 0/-!423!3]201 |CNLCCEF}TFGEBFT~F€FNJGNFMKFGPKVFMEFGWHCC€CENFLMNJF <=7>5?5@6A868 #0--<8A89^80/-00303-0-!]- 3213 3]7-39]7330 53735]3‚077<=7>5?5@6A86873<=7@[9\86871 !02343]32-203ƒ0 .23-3!]3/1-0-13   3]:0] 30-7%3!3x-; 0933#33-3003:9#;713 3]130 .]!-33053!1„#3 t„#7-5]3‚0!1„#3t„#73 -39]1-2-330 32323 .]-203-0:„#v„#; 303:…t;/<=7@[9\86830330/-0 0343]-0-Z03 -03-01-2  3]3-!1v„#3s#† $]356789=:"";--4 2.1 //--0‡9 !'/ !………3 6-ˆ]4"""7]-12-/213 3]0 3. „#s#3 ]03 %--:…t;-020/-0-0- 30] 7300/-31-2‰Š_?587‹[@\^>ŒA‹^@ :w].;&3 ]]Z 3--3/0-...  !03-0 3<=7@[9\868. 242 93!45!256789=:""…;71220/-0130 -0 3-4-33]] 31- 9- 300-56789=:"";021232130. 3305375]3‚07339]23 |CCEFIPGKKMDCBFGKF€CKVNHVŽJMIFGEBFCXNHVŽJMIFBXHMEWF 0 -0.243:$3/23$34"";0 2-0-723-3!]3/.<8A89^80 0213!-../012/ 31-2 20 20:x-;<8A89^80/33/2 v.230/]933#33-3003 :9#;730- -33]070]/230v.]/47130 !43.0:…tv;73! 0!41!0 +$/‘]-:……;.2421 -02 %20330]0<8A89^80- 3- 0-ƒ4 1-13 3]2-/130 33-203--4-20]-333 <8A89^80-0-!]-7-3305303- 03--43. …"7-5]3‚0 . "3-39]. v :%3!;#+']..:"";-0 <=7@[9\868-3301-2"03--4%2 3 --30/- 0//]-31-03--423 .]!4-]00]-07:";3//- 303:…t;3:…";-200]4 :%3!;%2033-... !03-0!4 3.:…tv;3$/‘]-+4!:""; 3-2//]/.32-2] !. <=7@[9\868-3301-22-203--4323 -!10 30-32-203--43]07 !/3]00 333<=7@[9\8681!0- 203 03 0:%3!; %2-0-!]-.<=7>5?5@6A8680/-! 2.0]2 3-:%3!;-3ƒ3 !2003 031-22!03-0 3 !49313-56789=:"";-ˆ3371220/-030 //]0-3300]!’/31-03--43-3- “XGH”KFGEBF•GNVECKF“HGEBCFKMNCKFGHCFME–XCEICKF|YF -03302312339]0- %2033/3Z3-22-203]0.3- 3!]3/.<8A89^870-0-5]3‚0 333053:x-;7/0--2-0 3 -Z-213/] 1-20]00-. 0- %2-0-. 3-333.//]/ <=7>5?5@6A8683<=7@[9\8687-23ƒ3 0- 0]-- 3---/323200/-0 234/23ƒ-/!23- %2 3!]30/-0.]2 1-2<8A89^807933#33-3003:9#; 1.]-3ƒ03 071—\6^?Œ\Š\9[@7 Œ\6Œ?8A^[@22!7—=7Œ\6Œ?8A^[@3\A8@@[@@7 :w--3 9 -2;1]07—?ŒA6˜Œ?5^@75[AŠ@6Œ‹8 #7—@65A^Œ?599Œ_@^@7™98\^89^@7:#303/3;3]`7 šŒ@\^?Œ›^@\[@7A8›^86[@722!`7š=7œ8^95@^^

#$%&'('!)'*+,-,./,1'! 

5&06(7123.456..,7./54-,./89&0 9%(7:;54<4;6/87,.,5,7=2$!>2$!7 ?-,<<,54@A4-,7436,;/3,72$!7B,C/3D.,7 @6;;,5,69E$ 'F7B64G6.@A/;6/87@6.,H/3,7I&J&77 B/5K83A/,7.4-6;K/,;,75)L77MG4;56..,7,D-/5, =9%!%>6&$7NOPQRSTUVWXYXZ[\PY5)L7 +8,<<4G/3524;7@,;GD-/]4-<67=5)L>^5#&7 _A/K484.6;/,7865/H6-, '$L77`a6.654;6<,7 H-65A,67b'659&7`c7@86DG43485,5D<7 #'6<6;Gcb'09&7dA,.,88/4;6<,7 ]-,D6;]6.G//7=5)L>I e0&&F7 dA,.,88/4;6<,782;6G-/]4-<67=5'>1&(7 dA,.,88/48/-,7,;HD.,5,7=5%>1&(7dc7G63/@/6;87 =5)L>25fg(7dc76336;5-/3,7=2$!> hF7dc7-45D.,7#)'&7dc78/<4;86;//1&( 0i%j?-,<<,54@A4-,7436,;/3,F7MG4;56..,7 ,D-/5,7&dA,.,88/48/-,(&&j&'E  L'$$(*)'f&&(L&(=9&L&'657,.cF "">7L$+c7]6;685-,5,L&((E'!

#Ek)'04!="">')$& +c78D.3,5,E&!)(&(&&''E&!E&)( lmnlonpqrsltunoltlvlwqxnryxnmpzon{|qoq|sqon}qttnlyn (' &%E(7!)'(%E$&'E&) $&( &&'&E'E&'&'( E 'E&0$'(()%7$&)$(!($& qysl|~yqymrtnv~yxlml~yonly€zqyvqnmpqn~|{p~t~un &&!)&E*+c78D.3,5,c7I$()(*)' $(()%($L$&+c7]6;685-,5,7E&&( !)('$'(%*()%' &'7!E&)(''( &!)&'(&&E('$()$&* $E)%‚$L'(7'J+c78D.3,5,FE* (' &&J'E(& (7$'&' rƒ~zmn{rtq~qysl|~yqymrtnv~yxlml~yonvrynƒqnxl„wvztm…n †$$&&&$'(' ozƒolxlqon„~|n„zmz|qnomzxlqon|qtrmqxnm~nmpqnly€zqyvqn *' &E''( $%& E%7$'E&'*%$&''E&'& *$((E'('E&(&' (

‡&(ˆ#$( 3&(5& 5)&‰( Š&9) ""

f&)&% I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŒhŽ I 9&''‹%‹ŽŒh i!)&% I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŽŒh I 9&''‹%‹ŒhŽ #&E$ I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŽŒh I 9&''‹%‹ŽŒh 6' I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŒhŽ 9&I ''‹%‹"ŽŒh #&% I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŽŒh I 9&''‹%‹ŒŽh f)% I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŽŒh 9&I ''‹%‹ŽŒh 6))( I ‹Œh

9&''%‹"Ž 9&I ''‹%‹"ŽŒh I 9&''‹%‹ŽŒh 9 ! I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŒhŽ I 9&''‹%‹ŒhŽ ‚E! I ‹"Œh

9&''%‹"Ž 9&I ''‹%‹"ŒhŽ I 9&''‹%‹"ŒŽh 4 ! I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŒhŽ I 9&''‹%‹ŽŒh ^E ! I ‹

9&''%‹Ž 9&I ''‹%‹ŒŽh I 9&''‹%‹ŽŒh ""

f&)&% I ‹Œh

9&''%‹Ž 9&I ''‹%‹ŒhŽ I 9&''‹%‹ŒhŽ

‘’“”•–—˜2' &&&'3&(5&75)&‰(&Š&9)('(79&& h&&'&0(&=9&&h&&'&9&7 &'>7!Lf&)&%""&f&)&%""

# 563$067%7&6'56&&07(79)*+6%)90%6447_{3,'6*(632990'-673}

6$.4)/'2(52%24&9

/012(+31 45'1!163$1 %37873129371:*;9$<=843834392% 11>88?6 37$41'4731)331=70371 84307 3787>1!1>?(@8$18 ABCDEFGGBHIDJHKLMNHGFCOMBEPQHRMGDHSLHTUFEVHFEHFEDEWCDXGH 3Y=784737 8

32;2324$29

ZXGLMFTD[\DEBDMMB]H\Q^QHH_`abQHHcBFTDCDdeJXMFHfZFgBMMFJBDOUWgLFLhH 1887311ijklmnopqrstuvwqnx5/4=<'1 44y 5300>71<31&873351947 1830033193 z3717p{ $7@,(4==73""0>127 43> 7>{373813391;17387 1>%13 27>889387|{| $1Y=73%}}&1y >1=48418474=0 1373731 ~owotopxrso€o@41=0>7>7 {""" ;73;1875,941%83 173%}}# (31 ‚71883  1410174!3ƒ ;3163879$7 18308417{#} ;73;18757941%83 173%,%31;307 1}}#(31 ‚71833&1187%14133 &1187;316387911$11317 1830$18 52y741818=1 31830383171$017171< 08417p{}"

517317%""~owotopntl€tso81381 37 31 = 911$1131917 13(31 381707 |{##|

5!„07$7/38037.176!747-1 482 7@22, 5875…††`QHH‡EdXLEgLHDIHEXTJBLETG]HTLCOLJFTXJL]HMBPUTH

18133>07747=~owotopxrso€o111 318740916ˆ4137 3>7‰{|} 1!87'6,%7ˆ437%3}}& 113138 1 123731 188 !18314!4 3 =0840913=5317 3380 $4 !317$11‚%1327>889387 "{# .3761;,%87@&"" 170 0>1 3781 OJDKMLHGLŠXLEgLHIDJHFHULFNMFENH‹FWHgDFGTQHŒDXJEFMHDIH^DFGTFMH 381707{|| %175331>7%',.17 1821705"<5373== 313Y3030~owotopvmtxltmop5 173130>1<8738 7 y 1>7}"{" %7ˆ437%3,4!7.""&031 ~owotop xrso€o1813 13371873837181 83 8284137$181190=97377{ }# %137797$Ž170$9,/870=@5""|<384 183 3188!1831 ‚718883 8 GXOLJKgBFBGHNDHIXENDHNFHFPXEFHNDGHFTDGQH‘JF’LM]Hbf_h“  %137797&17'$7$7'$,9187$ 5""}< 17%13&1888330$18113=33 51947 13{1> 01(31 237 @41=$1813817075<{# %417647 374)3317%$,41873 72$ "" 037 173130>1=01333841> 31 !47917 13 1331@41= 3>7 {}#" 3=876.}|#(383!431811313= 131 =~owotopxrso€o384=1783 8= 940 3380$4 !3108284137$1811 90=97377‰|{ ;148 ~”pntl€tso ~”punmnx€wo€o ~”pxrso€o (3 8385• < #–" "#531 < ""531 < 911311 /1238478 3 8  128 $4!11783=401=17 )3=801 13 $13=4 4!17801=17 (4!3=80 1 13 -1=17=037111 118314174>3183!438 01 13 3131338)3= 801 13 3138130381(4! 133= 2y1 1=03711>1 '1_11__388133 ;3813338 '1_1₃_48₁Y3_3_803_3!4837_3801 '323838 9141Y308 084=17 91_41Y3018310 _9141Y3018310 013 4114 Y03 =01 )53=0 13 3==0 191Y13n€po”7 ""< )03 =01 53= 13= 91Y13n€po”7 ""< 3=7 517315""< &03884> &03884>

#$%&'('!)'*+,-,./,1'! 

3(&70#123'& 45&( &'$&(()&'&( 6& &&773'5&9)7 &('80$''&719 :1:

9&;&'7<74&) 76=6%&7>""6$?&* +,-,./,@ &5'&'$%5&A&$5&(*B&& $%5'&7::91

9' &(('7BC71 &7#C76&&'73=#&D)( B64""4)'&'&'(&5&(&;&D)&'% *9&&C&&'&0(&7 &'E8 ' &(79 9' (731:6$'& &F*$0'&G5&

2?''*3HIJ#KL#KM(5$)(!'(( 2!'((73'$3 'EN19"

9)&#(' &73#=3(G''&76#112 3'& 45&(@ &5'&'$%5&A&5O'5&(3'!'P

&M&&Q#&)&3'6&&(7M'&R'(79C7 &('0()&7S9112

9)&#(' &73#7M75'M&(757T&)&(9'&7 3=M&(7TM113'& 45&(&'(' U((5'(5& &P&(5&&(&& &'$& 9C &('V()&792"

9)&#(' &73#7T&)&(9'&7376&&'73#= WXYZ[\]^_`^aYbcdeccfghheccijkcl\Ym^nco^pYc\]cq^]rk\st^c T&7M'&R'(79C7 &'0()&7:"9: 672273'!)7937CU&70C9=6 '&7T29 

"""15(' &''T&&(&(T&7 3'5&9)7 &'B)&*C&(&3(&5$7 922