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MÉTODOS PARA SEGMENTAÇÃO NUCLEAR E AVALIAÇÃO DA EXPRESSÃO GÊNICA EM IMAGENS SAGITAIS DE EMBRIÕES DE DROSOPHILA MELANOGASTER

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❯◆■❱❊❘❙■❉❆❉❊ ❋❊❉❊❘❆▲ ❉❊ ❯❇❊❘▲➶◆❉■❆ ❋❆❈❯▲❉❆❉❊ ❉❊ ❈❖▼P❯❚❆➬➹❖

P❘❖●❘❆▼❆ ❉❊ PÓ❙✲●❘❆❉❯❆➬➹❖ ❊▼ ❈■✃◆❈■❆ ❉❆ ❈❖▼P❯❚❆➬➹❖

▼➱❚❖❉❖❙ P❆❘❆ ❙❊●▼❊◆❚❆➬➹❖ ◆❯❈▲❊❆❘ ❊

❆❱❆▲■❆➬➹❖ ❉❆ ❊❳P❘❊❙❙➹❖ ●✃◆■❈❆ ❊▼ ■▼❆●❊◆❙

❙❆●■❚❆■❙ ❉❊ ❊▼❇❘■Õ❊❙ ❉❊ ❉❘❖❙❖P❍■▲❆ ▼❊▲❆◆❖●❆❙❚❊❘

❉❆◆■❊▲❆ ❏❯❙❚■◆■❆◆❖ ❉❊ ❙❖❯❙❆

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❯◆■❱❊❘❙■❉❆❉❊ ❋❊❉❊❘❆▲ ❉❊ ❯❇❊❘▲➶◆❉■❆ ❋❆❈❯▲❉❆❉❊ ❉❊ ❈❖▼P❯❚❆➬➹❖

P❘❖●❘❆▼❆ ❉❊ PÓ❙✲●❘❆❉❯❆➬➹❖ ❊▼ ❈■✃◆❈■❆ ❉❆ ❈❖▼P❯❚❆➬➹❖

❉❆◆■❊▲❆ ❏❯❙❚■◆■❆◆❖ ❉❊ ❙❖❯❙❆

▼➱❚❖❉❖❙ P❆❘❆ ❙❊●▼❊◆❚❆➬➹❖ ◆❯❈▲❊❆❘ ❊

❆❱❆▲■❆➬➹❖ ❉❆ ❊❳P❘❊❙❙➹❖ ●✃◆■❈❆ ❊▼ ■▼❆●❊◆❙

❙❆●■❚❆■❙ ❉❊ ❊▼❇❘■Õ❊❙ ❉❊ ❉❘❖❙❖P❍■▲❆ ▼❊▲❆◆❖●❆❙❚❊❘

❉✐ss❡rt❛çã♦ ❞❡ ▼❡str❛❞♦ ❛♣r❡s❡♥t❛❞❛ à ❋❛❝✉❧❞❛❞❡ ❞❡ ❈♦♠✲ ♣✉t❛çã♦ ❞❛ ❯♥✐✈❡rs✐❞❛❞❡ ❋❡❞❡r❛❧ ❞❡ ❯❜❡r❧â♥❞✐❛✱ ▼✐♥❛s ●❡✲ r❛✐s✱ ❝♦♠♦ ♣❛rt❡ ❞♦s r❡q✉✐s✐t♦s ❡①✐❣✐❞♦s ♣❛r❛ ♦❜t❡♥çã♦ ❞♦ tít✉❧♦ ❞❡ ▼❡str❡ ❡♠ ❈✐ê♥❝✐❛ ❞❛ ❈♦♠♣✉t❛çã♦✳

➪r❡❛ ❞❡ ❝♦♥❝❡♥tr❛çã♦✿ ❇❛♥❝♦ ❞❡ ❉❛❞♦s ❡ ■♠❛❣❡♥s✳

❖r✐❡♥t❛❞♦r✿

Pr♦❢✳ ❉r✳ ❇r✉♥♦ ❆✉❣✉st♦ ◆❛ss✐❢ ❚r❛✈❡♥ç♦❧♦

❈♦♦r✐❡♥t❛❞♦r✿

Pr♦❢✳ ❉r✳ ❋r❛♥❝✐s❝♦ ❏♦sé P❡r❡✐r❛ ▲♦♣❡s

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❯◆■❱❊❘❙■❉❆❉❊ ❋❊❉❊❘❆▲ ❉❊ ❯❇❊❘▲➶◆❉■❆ ❋❆❈❯▲❉❆❉❊ ❉❊ ❈❖▼P❯❚❆➬➹❖

P❘❖●❘❆▼❆ ❉❊ PÓ❙✲●❘❆❉❯❆➬➹❖ ❊▼ ❈■✃◆❈■❆ ❉❆ ❈❖▼P❯❚❆➬➹❖

❖s ❛❜❛✐①♦ ❛ss✐♥❛❞♦s✱ ♣♦r ♠❡✐♦ ❞❡st❡✱ ❝❡rt✐✜❝❛♠ q✉❡ ❧❡r❛♠ ❡ r❡❝♦♠❡♥❞❛♠ ♣❛r❛ ❛ ❋❛❝✉❧✲ ❞❛❞❡ ❞❡ ❈♦♠♣✉t❛çã♦ ❛ ❛❝❡✐t❛çã♦ ❞❛ ❞✐ss❡rt❛çã♦ ✐♥t✐t✉❧❛❞❛ ✏▼ét♦❞♦s ♣❛r❛ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r ❡ ❛✈❛❧✐❛çã♦ ❞❛ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❡♠ ✐♠❛❣❡♥s s❛❣✐t❛✐s ❞❡ ❡♠❜r✐õ❡s ❞❡ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✑ ♣♦r ❉❛♥✐❡❧❛ ❏✉st✐♥✐❛♥♦ ❞❡ ❙♦✉s❛ ❝♦♠♦ ♣❛rt❡ ❞♦s r❡q✉✐s✐t♦s ❡①✐❣✐❞♦s ♣❛r❛ ❛ ♦❜t❡♥çã♦ ❞♦ tít✉❧♦ ❞❡ ▼❡str❡ ❡♠ ❈✐ê♥❝✐❛ ❞❛ ❈♦♠♣✉t❛çã♦✳

❯❜❡r❧â♥❞✐❛✱ ✷✾ ❞❡ ◆♦✈❡♠❜r♦ ❞❡ ✷✵✶✸

❖r✐❡♥t❛❞♦r✿

Pr♦❢✳ ❉r✳ ❇r✉♥♦ ❆✉❣✉st♦ ◆❛ss✐❢ ❚r❛✈❡♥ç♦❧♦ ❯♥✐✈❡rs✐❞❛❞❡ ❋❡❞❡r❛❧ ❞❡ ❯❜❡r❧â♥❞✐❛

❈♦♦r✐❡♥t❛❞♦r✿

Pr♦❢✳ ❉r✳ ❋r❛♥❝✐s❝♦ ❏♦sé P❡r❡✐r❛ ▲♦♣❡s ❯♥✐✈❡rs✐❞❛❞❡ ❋❡❞❡r❛❧ ❞♦ ❘✐♦ ❞❡ ❏❛♥❡✐r♦

❇❛♥❝❛ ❊①❛♠✐♥❛❞♦r❛✿

Pr♦❢✳ ❉r✳ ❏♦❛q✉✐♠ ❈❡③❛r ❋❡❧✐♣❡ ❯♥✐✈❡rs✐❞❛❞❡ ❞❡ ❙ã♦ P❛✉❧♦

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❯◆■❱❊❘❙■❉❆❉❊ ❋❊❉❊❘❆▲ ❉❊ ❯❇❊❘▲➶◆❉■❆ ❋❆❈❯▲❉❆❉❊ ❉❊ ❈❖▼P❯❚❆➬➹❖

P❘❖●❘❆▼❆ ❉❊ PÓ❙✲●❘❆❉❯❆➬➹❖ ❊▼ ❈■✃◆❈■❆ ❉❆ ❈❖▼P❯❚❆➬➹❖

❉❛t❛✿ ◆♦✈❡♠❜r♦ ❞❡ ✷✵✶✸

❆✉t♦r✿ ❉❛♥✐❡❧❛ ❏✉st✐♥✐❛♥♦ ❞❡ ❙♦✉s❛

❚ít✉❧♦✿ ▼ét♦❞♦s ♣❛r❛ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r ❡ ❛✈❛❧✐❛çã♦ ❞❛ ❡①♣r❡ssã♦ ❣ê✲ ♥✐❝❛ ❡♠ ✐♠❛❣❡♥s s❛❣✐t❛✐s ❞❡ ❡♠❜r✐õ❡s ❞❡ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r ❋❛❝✉❧❞❛❞❡✿ ❋❛❝✉❧❞❛❞❡ ❞❡ ❈♦♠♣✉t❛çã♦

●r❛✉✿ ▼❡str❛❞♦

❋✐❝❛ ❣❛r❛♥t✐❞♦ à ❯♥✐✈❡rs✐❞❛❞❡ ❋❡❞❡r❛❧ ❞❡ ❯❜❡r❧â♥❞✐❛ ♦ ❞✐r❡✐t♦ ❞❡ ❝✐r❝✉❧❛çã♦ ❡ ✐♠♣r❡ssã♦ ❞❡ ❝ó♣✐❛s ❞❡st❡ ❞♦❝✉♠❡♥t♦ ♣❛r❛ ♣r♦♣ós✐t♦s ❡①❝❧✉s✐✈❛♠❡♥t❡ ❛❝❛❞ê♠✐❝♦s✱ ❞❡s❞❡ q✉❡ ♦ ❛✉t♦r s❡❥❛ ❞❡✈✐❞❛♠❡♥t❡ ✐♥❢♦r♠❛❞♦✳

❆✉t♦r

❖ ❆❯❚❖❘ ❘❊❙❊❘❱❆ P❆❘❆ ❙■ ◗❯❆▲◗❯❊❘ ❖❯❚❘❖ ❉■❘❊■❚❖ ❉❊ P❯❇▲■❈❆➬➹❖ ❉❊❙❚❊ ❉❖❈❯▼❊◆❚❖✱ ◆➹❖ P❖❉❊◆❉❖ ❖ ▼❊❙▼❖ ❙❊❘ ■▼P❘❊❙❙❖ ❖❯ ❘❊P❘❖✲ ❉❯❩■❉❖✱ ❙❊❏❆ ◆❆ ❚❖❚❆▲■❉❆❉❊ ❖❯ ❊▼ P❆❘❚❊❙✱ ❙❊▼ ❆ P❊❘▼■❙❙➹❖ ❊❙❈❘■❚❆ ❉❖ ❆❯❚❖❘✳

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❉❡❞✐❝❛tór✐❛

❆ ❉❡✉s P❛✐✱ ❆ ▼❛r✐❛ ❙❛♥tíss✐♠❛ ▼ã❡✱

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❆❣r❛❞❡❝✐♠❡♥t♦s

Pr✐♠❡✐r❛♠❡♥t❡ ❛ ❉❡✉s✱ ❙❡♥❤♦r ❞❡ ♠✐♥❤❛ ❱✐❞❛✱ ♣♦r ♣❡r♠✐t✐r✲♠❡ s♦♥❤❛r ❡ ❧✉t❛r ♣❡❧❛ r❡❛❧✐③❛çã♦ ❞❡ss❛ ✐♠♣♦rt❛♥t❡ ❝♦♥q✉✐st❛✳ ❖❜r✐❣❛❞❛ P❛✐ ❛♠❛❞♦✱ ♣♦r ❝✉✐❞❛r ❞❡ ❝❛❞❛ ❞❡t❛✲ ❧❤❡✱ ♣♦r ❝♦❧♦❝❛r ♣❡ss♦❛s ❛❜❡♥ç♦❛❞❛s ❛♦ ♠❡✉ r❡❞♦r ❡ ❢♦rt❛❧❡❝❡r✲♠❡ ♥♦s ♠♦♠❡♥t♦s ❞✐❢í❝❡✐s✳ ❖❜r✐❣❛❞❛ ♣❡❧❛ ❜❡♥çã♦ ♣r❡st❛❞❛ ♥❛ ❝♦♥❝❧✉sã♦ ❞❡ ♠❛✐s ✉♠ ❝✐❝❧♦ ✐♠♣♦rt❛♥t❡ ❡♠ ♠✐♥❤❛ ✈✐❞❛✱ ♣❡❧❛ ♠✐s❡r✐❝ór❞✐❛ ❡ ❛♠♦r ✐♥✜♥✐t♦✳ ❆ ❚✐ ♠❡✉ ❙❡♥❤♦r t♦❞❛ ❤♦♥r❛ ❡ ❣❧ór✐❛✦

❆♦ ♠❡✉ ❡st✐♠❛❞♦ ♦r✐❡♥t❛❞♦r✲♣r♦❢❡ss♦r ❇r✉♥♦ ❆✉❣✉st♦ ◆❛ss✐❢ ❚r❛✈❡♥ç♦❧♦✱ ❛❣r❛❞❡ç♦ ❛ ♣❛❝✐ê♥❝✐❛✱ ❞❡❞✐❝❛çã♦ ❡ ♦r✐❡♥t❛çã♦ ❝♦♥❝❡❜✐❞❛✳ ❯♠ sá❜✐♦ t✉t♦r✱ ♦ r❡s♣❡✐t♦ ❡ ❝♦♥✜❛♥ç❛ ❡♠ ♠✐♠ ❞❡♣♦s✐t❛❞♦s✱ ❝♦♠ ❝❡rt❡③❛✱ ❢♦r❛♠ ❡ss❡♥❝✐❛✐s ♣❛r❛ ❛ ❝♦♥❝❧✉sã♦ ❞❡ss❛ ♣❡sq✉✐s❛✳ ❖❜r✐❣❛❞❛ ♣❡❧❛ s❡r❡♥✐❞❛❞❡ ❡ s❡❣✉r❛♥ç❛ ❞✐s♣❡♥s❛❞❛ ❡♠ ♠❡✉s ♠♦♠❡♥t♦s ❞❡ ❛♥s✐❡❞❛❞❡ ❡ ❞ú✈✐❞❛s✳ ❉❡ t✐ ❧❡✈❛r❡✐ ♠✉✐t♦ ❛❧é♠ ❞♦s ❡♥s✐♥❛♠❡♥t♦s ❝✐❡♥tí✜❝♦s✱ ✜❝❛ ❛ ❛❞♠✐r❛çã♦ ♣❡❧♦ s❡✉ ♣r♦✜ss✐♦♥❛❧✐s♠♦✱ ❜♦♥❞❛❞❡ ❡ s❛❜❡❞♦r✐❛✳

❆♦ ❝♦♦r✐❡♥t❛❞♦r ❋r❛♥❝✐s❝♦ ❏♦sé P❡r❡✐r❛ ▲♦♣❡s✱ ❛❣r❛❞❡ç♦ ❛ ❛t❡♥çã♦ ❞✉r❛♥t❡ ❛ ❝♦♥str✉çã♦ ❞❡ss❡ tr❛❜❛❧❤♦✳ ❖❜r✐❣❛❞❛ ♣❡❧❛s ✐♠❛❣❡♥s ❝♦♥❝❡❞✐❞❛s ❡ ♣❡❧❛ ❞✐s♣♦♥✐❜✐❧✐❞❛❞❡ ❡♠ ❞✐s❝✉t✐✲❧❛s✳ ❆♦ ♠❡✉ ❡t❡r♥♦ ♠❡str❡ ❆♥❞ré ❆✉❣✉st♦ ❈❛❧❛❜r❡③✱ ♣♦r ❛♥♦s ❛trás ✐♥❝❡♥t✐✈❛r✲♠❡ ❛ ❝♦✲ ♥❤❡❝❡r ❛ ♣❡sq✉✐s❛✳ ▼♦t✐✈❛♥❞♦✲♠❡ ❛ tr✐❧❤❛r ♦ ❝❛♠✐♥❤♦ ♣♦r ♦♥❞❡ t❡♥❤♦ ❛♥❞❛❞♦✳

❆ ✈♦❝ê ♠✐♥❤❛ ♠ã❡✱ ♦ ❛♠♦r ♠❛✐s s✉❜❧✐♠❡ ❡ ♣✉r♦ q✉❡ ❉❡✉s ♣♦❞❡r✐❛ ♠❡ ❞❛r✳ ▼✐♥❤❛ r❡❢❡rê♥❝✐❛ ❡ ♠❡✉ ❡①❡♠♣❧♦ ❞❡ ✈✐❞❛✳ ❖❜r✐❣❛❞❛ ♣♦r ✐❧✉♠✐♥❛r ♦s ♠❡✉s ❞✐❛s ❝♦♠ t✉❛s ❞♦❝❡s ♣❛❧❛✈r❛s ❡ t❡✉ ❧✐♥❞♦ s♦rr✐s♦✳ ❖❜r✐❣❛❞❛ ♣♦r s❡♠♣r❡ ❛❝r❡❞✐t❛r ❡♠ ♠✐♠✱ ♠❡s♠♦ q✉❛♥❞♦✱ às ✈❡③❡s✱ ♥❡♠ ❡✉ ♠❡s♠❛ ❛❝r❡❞✐t❛✈❛✳ ❖❜r✐❣❛❞❛ ♣❡❧♦ ❛♣♦✐♦ ♥♦s ♠♦♠❡♥t♦s ❞✐❢í❝❡✐s ❡ ❞❡ ❝❛♥s❛ç♦✳ ❖❜r✐❣❛❞❛ ♣❡❧❛ ♣r❡s❡♥ç❛ ❡ ✈✐❜r❛çã♦ ❡♠ ❝❛❞❛ ✈✐tór✐❛✳ ▼✐♥❤❛ ❡t❡r♥❛ ❣r❛t✐❞ã♦ ♣❡❧♦ t❡✉ ❛♠♦r✱ ✐♥❝❡♥t✐✈♦ ❡ ❝♦♥✜❛♥ç❛ ❛❜s♦❧✉t❛✱ ❝♦♠ ❝❡rt❡③❛ ❛s ❜❛s❡s q✉❡ ❡✉ ♣r❡❝✐s❛✈❛ ♣❛r❛ ✈❡♥❝❡r✳ ❊ss❛ ❝♦♥q✉✐st❛ é ♣♦r ✈♦❝ê✦

❆ ✈♦❝ê ♠❡✉ ❛♠❛❞♦ ❡s♣♦s♦✱ ❛❣r❛❞❡ç♦ ♦ ❛♠♦r✱ ❞❡❞✐❝❛çã♦ ❡ ❝♦♠♣r❡❡♥sã♦ ✐♥❝♦♥❞✐❝✐♦♥❛❧✳ P♦r s❡r ♠❡✉ ✜❡❧ ❝♦♠♣❛♥❤❡✐r♦✱ ♠❡s♠♦ ♥❛s ♠❛❞r✉❣❛❞❛s ❡♠ ❝❧❛r♦✳ ❆❣r❛❞❡ç♦✲t❡ ♣♦r ❝✉✐❞❛r tã♦ ❜❡♠ ❞❡ ♠✐♠✱ ♣❡❧♦ ❛♣♦✐♦ ❡♠♦❝✐♦♥❛❧✱ ❡ ♣♦r t♦❞♦s ♦s ❣❡st♦s ❞❡ ❛t❡♥çã♦ ❡ ❝❛r✐♥❤♦✳ P♦r t❡r t♦❧❡r❛❞♦ ♦s ❞✐❛s ❡ ♥♦✐t❡s ❞❡ ❡st✉❞♦ ❡♠ q✉❡ ♠❡ s❡♣❛r❡✐ ❞❡ t✐✱ ♣♦r t❡r ❡s♣❡r❛❞♦ ❡ ♠❡ ❛♠❛❞♦✳ ▼❡✉ ❛♠♦r✱ s✉❛ ♣r❡s❡♥ç❛ tr❛③ ❧✉③ ❡ s❡r❡♥✐❞❛❞❡ ♣❛r❛ ♦s ♠❡✉s ❞✐❛s✳ ❚❡ ❛♠♦✦

❆♦s ♠❡✉s ✐r♠ã♦s ❊❞♥❡② ❡ ❏❛❝q✉❡❧✐♥❡✱ ❧✐♥❞♦s ♣r❡s❡♥t❡s ❞❡ ❉❡✉s ❡♠ ♠✐♥❤❛ ✈✐❞❛✳ ❆❣r❛✲ ❞❡ç♦ ♦ ❛♠♦r✱ ❝❛r✐♥❤♦ ❡ ❛♣♦✐♦✳ ❖❜r✐❣❛❞❛ ♣♦r ❝♦♠♣r❡❡♥❞❡r❡♠ ♠✐♥❤❛ ❛✉sê♥❝✐❛✱ ♣❡❧❛ ♣❛✲ ❝✐ê♥❝✐❛ ❡ ✐♥❝❡♥t✐✈♦ ❝♦♥st❛♥t❡✳ ◆ã♦ ✐♠❛❣✐♥♦ ♠✐♥❤❛ ✈✐❞❛ s❡♠ ✈♦❝ês✳ ❆ ✈♦❝ê ♠✐♥❤❛ ✐r♠ã✱ ❡♠ ❡s♣❡❝✐❛❧✱ ❛❣r❛❞❡ç♦ ♣❡❧♦s três ❣r❛♥❞❡s ❛♠♦r❡s q✉❡ tr♦✉①❡ às ♥♦ss❛s ✈✐❞❛s✱ ♠❡✉s ❛♠❛❞♦s s♦❜r✐♥❤♦s✳ ❈♦♠ t♦❞❛ ❝❡rt❡③❛✱ ♠✐♥❤❛ ❢♦♥t❡ ❞❡ ✐♥s♣✐r❛çã♦✳ ▼❛t❡r✐❛❧✐③❛çã♦ ❞♦ ❛♠♦r ❞❡ ❉❡✉s ❡♠ ♠✐♥❤❛ ✈✐❞❛✳

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■❧③❛ ❡ ❈✐❞❛✱ ♦❜r✐❣❛❞❛ ♣♦r ❛❝♦♠♣❛♥❤❛r❡♠ ♠✐♥❤❛ tr❛❥❡tór✐❛ ❡ s❡r❡♠ ♣r❡s❡♥ç❛ ❢✉♥❞❛♠❡♥t❛❧ ❡♠ ♠✐♥❤❛ ✈✐❞❛✳ ❆❣r❛❞❡ç♦✲❧❤❡s ♣❡❧♦s ❛❜r❛ç♦s ❡ ❝❛r✐♥❤♦ ❛ ❝❛❞❛ r❡t♦r♥♦ ❛♦ s❡✐♦ ❢❛♠✐❧✐❛r✳

➚s ♠✐♥❤❛s q✉❡r✐❞❛s ❛✈ós✱ ❉✐❝❛ ❡ ▲♦✉r❞❡s ✭s❛✉❞❛❞❡s✮✱ ♣♦r t❡r❡♠ s✐❞♦ ❛s ♣r✐♠❡✐r❛s ♣❡ss♦❛s ❛ ♠❡ ✐♥❝❡♥t✐✈❛r✱ ♠♦t✐✈❛♥❞♦✲♠❡ ❛ ❝♦❧♦❝❛r ❡♠ ♣rát✐❝❛ ✈❛❧♦r❡s ✐♠♣♦rt❛♥t❡s ♣❛r❛ q✉❡ ❡✉ ❝❤❡❣❛ss❡ ❛té ❛q✉✐✱ ❝♦♠♦ ❛ ♣❡rs✐stê♥❝✐❛✱ ❢é ❡ ❛ ❞❡t❡r♠✐♥❛çã♦✳ ❆ ♠✐♥❤❛ ❛♠❛❞❛ ❱ó ◆❛✐r✱ ♣❡❧❛s ♦r❛çõ❡s ❡ ❝❛r✐♥❤♦✱ q✉❡ ❛♣❡s❛r ❞❛ ❞✐stâ♥❝✐❛ s❡♠♣r❡ ❢♦r❛♠ ❝♦♥st❛♥t❡s✳

❆♦s ♠❡✉s q✉❡r✐❞♦s ❛♠✐❣♦s✱ ❞❡ ❧♦♥❣❡ ❡ ❞❡ ♣❡rt♦✱ ❛ ✈♦❝ês✱ ♦ ♠❡✉ s✐♥❝❡r♦ ❛❣r❛❞❡❝✐♠❡♥t♦ ♣❡❧♦ ❝❛r✐♥❤♦✱ ❛♠✐③❛❞❡ ❡ t♦r❝✐❞❛✳ ❊♠ ❡s♣❡❝✐❛❧✱ às ♠✐♥❤❛s ✐r♠ãs ❞❡ ❝♦r❛çã♦✱ ❛♥❥♦s ❡♥✈✐❛❞♦s ♣♦r ❉❡✉s ♣❛r❛ ✐❧✉♠✐♥❛r ❛ ♠✐♥❤❛ ✈✐❞❛✳ ❆ ✈♦❝ês ◆❛②❛r❛✱ P♦❧❧②❛♥❛✱ ❉❛♥✐❡❧❛ ❡ ❋❧á✈✐❛✱ ❛❣r❛❞❡ç♦ ♦ ✐♥❝❡♥t✐✈♦✱ ♠❡s♠♦ q✉❡ ❞✐st❛♥t❡✳ ❖❜r✐❣❛❞❛ ♣❡❧❛s ❧♦♥❣❛s ❡ ❝✉rt❛s ❝♦♥✈❡rs❛s✱ ♣❡❧♦s ❝♦♥s❡❧❤♦s✱ r✐s♦s✱ ❝❤♦r♦s ❡ ❛❧❡❣r✐❛s✳

❆♦ ♠❡✉ ❛♠✐❣♦ ❘♦♥❞✐♥❡❧❡✱ q✉❡ ❝♦♠♣❛rt✐❧❤♦✉ ❝♦♠✐❣♦ ❝♦♥q✉✐st❛s✱ ❡s♣❛ç♦s✱ s❛✉❞❛❞❡s✱ s♦♥❤♦s✱ ❡ ❛❧❣✉♥s ❞❡s❛✜♦s✳ ❆❣r❛❞❡ç♦ ❛ ❛♠✐③❛❞❡ ❡ ♦ ❛♣♦✐♦ ❝♦♥st❛♥t❡✱ ❡ ❡s♣❡❝✐❛❧♠❡♥t❡✱ s✉❛ ❛❧❡❣r✐❛✳

➚ ♠✐♥❤❛ ❝♦♠♣❛♥❤❡✐r❛ ❞❡ ❥♦r♥❛❞❛ ❛❝❛❞ê♠✐❝❛✱ ❱✐✈✐❛♥❡✳ ❙♦✉ ❣r❛t❛ ♣❡❧❛ ❛♠✐③❛❞❡✱ t♦r❝✐❞❛ ❡ ❝❛r✐♥❤♦✳ ❋♦✐ ♠❛r❛✈✐❧❤♦s♦ ❞✐✈✐❞✐r ♠❛✐s ✉♠❛ ❡①♣❡r✐ê♥❝✐❛ ❝♦♠ ✈♦❝ê✳

❆♦s ♠❡✉s ❝♦❧❡❣❛s ❞♦ ♠❡str❛❞♦ ❡ ❞♦ ●r✉♣♦ ❞❡ P❡sq✉✐s❛ ❇❛♥❝♦ ❞❡ ❉❛❞♦s ❡ ■♠❛❣❡♥s✱ ❛❣r❛❞❡ç♦ ♦ ❝♦♥✈í✈✐♦ ❤❛r♠♦♥✐♦s♦ ❡ r❡s♣❡✐t♦✳ ❱♦✉ ❧❡✈á✲❧♦s ♣❛r❛ s❡♠♣r❡ ❡♠ ♠❡✉ ❝♦r❛çã♦✳ ❉❡ ♠♦❞♦ ❡s♣❡❝✐❛❧✱ ❛❣r❛❞❡ç♦ ❛ ❈❧❡✐❛♥❡✱ ❏✉❧✐❡t❡ ❡ ❈í❝❡r♦✱ ♣❡❧❛ ❛♠✐③❛❞❡ ❡ ❡stí♠✉❧♦s ❞❡❞✐❝❛❞♦s ❞✉r❛♥t❡ ❡ss❛ ❢❛s❡✳

❆ t♦❞♦s ♦s ❛♠✐❣♦s ❞♦ ▼❯❘✱ ●PP ❡ ❊❈❈✱ ❞♦s q✉❛✐s s❡♠♣r❡ ♣✉❞❡ ❝♦♥t❛r ❝♦♠ s✉❛s ♦r❛çõ❡s ❡ ♣r❡st❡③❛✳ ❍♦❥❡ sã♦ ♣❛rt❡ ❞❡ ♠✐♥❤❛ ❢❛♠í❧✐❛✱ ❢♦♥t❡ ❞❡ ❧✉③ ❡ ❢r❛t❡r♥✐❞❛❞❡ ❡♠ ♠✐♥❤❛ ✈✐❞❛✳

❆♦ ❝♦r♣♦ ❞♦❝❡♥t❡ ❞❛ ❋❆❈❖▼✱ ♣❡❧♦s ❡♥s✐♥❛♠❡♥t♦s ❛❝❛❞ê♠✐❝♦s ❢✉♥❞❛♠❡♥t❛✐s ♣❛r❛ ❛ ❝♦♥❝❧✉sã♦ ❞❡ss❡ tr❛❜❛❧❤♦✳

❆♦ ❈◆Pq ♣❡❧❛ ❝♦♥❝❡ssã♦ ❞❛ ❜♦❧s❛ ❞✉r❛♥t❡ ♦ ❞❡s❡♥✈♦❧✈✐♠❡♥t♦ ❞❛ ♣❡sq✉✐s❛✳

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❘❡s✉♠♦

◆♦s ú❧t✐♠♦s ❛♥♦s✱ ❛s té❝♥✐❝❛s ❞❡ ♣r♦❝❡ss❛♠❡♥t♦ ❞✐❣✐t❛❧ ❞❡ ✐♠❛❣❡♥s tê♠ s✐❞♦ ❧❛r❣❛✲ ♠❡♥t❡ ✉t✐❧✐③❛❞❛s ❡♠ ❝♦♥❥✉♥t♦ ❝♦♠ ♦✉tr❛s ár❡❛s ❞❛ ❝✐ê♥❝✐❛✱ ✐♥❝❧✉✐♥❞♦ ❛ ❜✐♦❧♦❣✐❛✳ ❊st❛s té❝♥✐❝❛s ❣❡r❛❧♠❡♥t❡ ✈✐s❛♠ ♦t✐♠✐③❛r ♦ t❡♠♣♦✱ ❛✉♠❡♥t❛r ❛ ♣r❡❝✐sã♦ ❡ ❡❧✐♠✐♥❛r ❛ s✉❜❥❡t✐✈✐✲ ❞❛❞❡ ❞♦ ♣r♦❝❡ss♦ ❞❛ s✐♠♣❧❡s ❛♥á❧✐s❡ ✈✐s✉❛❧✳ ❊ss❡ ❝r❡s❝✐♠❡♥t♦ ♠♦t✐✈♦✉✲s❡✱ ♣r✐♥❝✐♣❛❧♠❡♥t❡✱ ♣♦r ❛✈❛♥ç♦s ✐♠♣♦rt❛♥t❡s ♥❛ ❇✐♦❧♦❣✐❛ ❈♦♠♣✉t❛❝✐♦♥❛❧ ❡ ♣❡❧♦ ❛✉♠❡♥t♦ ❝♦♥s✐❞❡rá✈❡❧ ❞♦s ❜❛♥✲ ❝♦s ❞❡ ✐♠❛❣❡♥s ❜✐♦❧ó❣✐❝❛s✳ ❖ ♣r❡s❡♥t❡ tr❛❜❛❧❤♦ ♣r♦♣õ❡ ♠ét♦❞♦s ♣❛r❛ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r ❡ ❛✈❛❧✐❛çã♦ ❞❛ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❡♠ ✐♠❛❣❡♥s ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ❖ ♠ét♦❞♦ ❞❡ s❡❣♠❡♥t❛çã♦ ♣r♦♣♦st♦ ❛❜♦r❞❛ ❞✐❢❡r❡♥t❡s té❝♥✐❝❛s ❞❡ ♣r♦❝❡ss❛♠❡♥t♦ ❞✐❣✐t❛❧ ❞❡ ✐♠❛❣❡♥s✱ ❡ ❝♦♥s✐st❡ ❡♠ r❡❛❧✐③❛r ❛ s❡❣♠❡♥t❛çã♦ ❛✉t♦♠át✐❝❛ ❞♦s ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s ❝♦♥t✐❞♦s ❡♠ ✐♠❛✲ ❣❡♥s s❛❣✐t❛✐s ❞❡ ❡♠❜r✐õ❡s ❞❡ss❡ ♠♦❞❡❧♦ ❜✐♦❧ó❣✐❝♦✳ ❖ ♠ét♦❞♦ ❜❛s❡✐❛✲s❡ ♥❛ ❡st✐♠❛t✐✈❛ ❞❡ ❝✉r✈❛t✉r❛ ❞♦ ❝♦♥t♦r♥♦ ❞♦s ♦❜❥❡t♦s✱ ❡ ❝♦♥s✐st❡ ❡♠ s❡♣❛r❛r ♥ú❝❧❡♦s ❛❣r❡❣❛❞♦s ♣♦r ♠❡✐♦ ❞❡ s❡❣♠❡♥t♦s tr❛ç❛❞♦s ❡♥tr❡ ♣♦♥t♦s ❝ô♥❝❛✈♦s ❝♦rr❡s♣♦♥❞❡♥t❡s✳ ❆ ❛❜♦r❞❛❣❡♠ ❛♣r❡s❡♥t❛❞❛ é ♥♦✈❛ ♣❛r❛ ❡ss❡ ❝❡♥ár✐♦✱ ❡ ❝♦♥tr✐❜✉✐ ♣❛r❛ ❛ s♦❧✉çã♦ ❞♦ ♣r♦❜❧❡♠❛ ❞❡ ❝♦♠♣♦♥❡♥t❡s ❛❣r❡❣❛❞♦s✱ ❝❛r❛❝t❡ríst✐❝❛ r❡❝♦rr❡♥t❡ ❡♠ ❛♣❧✐❝❛çõ❡s ✈♦❧t❛❞❛s à s❡❣♠❡♥t❛çã♦ ❞❡ ♥ú❝❧❡♦s✳ ❖s ♠ét♦❞♦s ❞❡ ❛✈❛❧✐❛çã♦ ✐♥❝❧✉❡♠ ❛❜♦r❞❛❣❡♥s ♣❛r❛ q✉❛♥t✐✜❝❛çã♦ ❡ ✈✐s✉❛❧✐③❛çã♦ ❞❛ ✐♥t❡♥s✐❞❛❞❡ ♠é❞✐❛ ❞❡ ♣r♦t❡í♥❛ ❡ ❘◆❆ ♥❛s r❡❣✐õ❡s ✐♥tr❛♥✉❝❧❡❛r ❡ ❝✐t♦♣❧❛s♠át✐❝❛ ❞♦s ❡♠❜r✐õ❡s ❞❡ ❉r♦s♦♣❤✐❧❛✳ ❖ ♠ét♦❞♦ ❞❡ q✉❛♥t✐✜❝❛çã♦✱ ❡♠ ❡s♣❡❝í✜❝♦✱ ♣♦ss✐❜✐❧✐t❛ ♦❜t❡r ♦ ♣❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❛ss✉♠✐❞♦ ❛♦ ❧♦♥❣♦ ❞♦ ❡✐①♦ ❛♥t❡r✐♦r✲♣♦st❡r✐♦r ❞♦s ❡♠❜r✐õ❡s ❞❡ ❉r♦s♦♣❤✐❧❛✳ P♦r ♦✉tr♦ ❧❛❞♦✱ ♦ ♠ét♦❞♦ ❞❡ ✈✐s✉❛❧✐③❛çã♦ ✐♥❝❧✉✐ ✉♠❛ ♥♦✈❛ ❛❜♦r❞❛❣❡♠ ♣❛r❛ ❡①✐❜✐çã♦ ❞❛s ✐♠❛❣❡♥s s❛❣✐t❛✐s✱ ❝❛♣❛③ ❞❡ ♦r❣❛♥✐③❛r ♦s ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s ❡♠ ✉♠❛ ❝♦♥✜❣✉r❛çã♦ ❧✐♥❡❛r✳ ❖ ✉s♦ ❡✜❝✐❡♥t❡s ❞♦s ♠ét♦❞♦s ❞❡ ❛✈❛❧✐❛çã♦ ❛✉①✐❧✐❛ ❛ ✐♥t❡r♣r❡t❛çã♦ ❞♦s ❞❛❞♦s ♦❜t✐❞♦s✳ ❊♠ ❣❡r❛❧✱ ♦s ♠ét♦❞♦s ♣❛r❛ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r ❡ ❛✈❛❧✐❛çã♦ ❞❛ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ r❡♣r❡s❡♥t❛♠ ✉♠ ♣❛ss♦ ❡ss❡♥✲ ❝✐❛❧ ♣❛r❛ ❛ r❡❛❧✐③❛çã♦ ❞❡ ❞✐✈❡rs❛s ❛♥á❧✐s❡s ❜✐♦❧ó❣✐❝❛s s♦❜r❡ ❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ❊①♣❡r✐♠❡♥t♦s ❢♦r❛♠ r❡❛❧✐③❛❞♦s ❛ ✜♠ ❞❡ ✐❧✉str❛r ♦ ❞❡s❡♠♣❡♥❤♦ ❞❛ s❡❣♠❡♥t❛çã♦✱ q✉❛♥❞♦ ❝♦♠♣❛r❛❞♦ ❝♦♠ ♦✉tr❛s ❛❜♦r❞❛❣❡♥s ❝♦♠✉♠❡♥t❡ ❡♥❝♦♥tr❛❞❛s ♥❛ ❧✐t❡r❛t✉r❛✳ ❖s r❡s✉❧t❛❞♦s ♦❜t✐❞♦s ❞❡♠♦♥str❛r❛♠ q✉❡ ♦ ♠ét♦❞♦ ♣r♦♣♦st♦ é ❡✜❝❛③ ❛ ♥í✈❡❧ q✉❛❧✐t❛t✐✈♦ ❡ q✉❛♥t✐t❛t✐✈♦✳ ❖ ♠ét♦❞♦ ♣♦❞❡ ❛✐♥❞❛ s❡r ✉t✐❧✐③❛❞♦ ❡♠ ❞✐❢❡r❡♥t❡s ❛♣❧✐❝❛çõ❡s ❡ ár❡❛s✱ t❛✐s q✉❡ ❡♥✈♦❧✈❛♠ ❝♦♥✜❣✉r❛çõ❡s s✐♠✐❧❛r❡s✳

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(17)

❆❜str❛❝t

■♥ r❡❝❡♥t ②❡❛rs✱ t❤❡ t❡❝❤♥✐q✉❡s ♦❢ ❞✐❣✐t❛❧ ✐♠❛❣❡ ♣r♦❝❡ss✐♥❣ ❤❛✈❡ ❜❡❡♥ ✇✐❞❡❧② ✉s❡❞ ✐♥ ❝♦♥❥✉♥❝t✐♦♥ ✇✐t❤ ♦t❤❡r ❛r❡❛s ♦❢ s❝✐❡♥❝❡✱ ✐♥❝❧✉❞✐♥❣ ❜✐♦❧♦❣②✳ ❚❤❡s❡ t❡❝❤♥✐q✉❡s ❣❡♥❡r❛❧❧② ❛✐♠ t♦ ♦♣t✐♠✐③❡ t✐♠❡✱ ✐♥❝r❡❛s❡ ❛❝❝✉r❛❝② ❛♥❞ ❡❧✐♠✐♥❛t❡ t❤❡ s✉❜❥❡❝t✐✈✐t② ♦❢ t❤❡ ♣r♦❝❡ss ♦❢ s✐♠♣❧❡ ✈✐s✉❛❧ ❛♥❛❧②s✐s✳ ❚❤✐s ✐♥❝r❡❛s❡ ✇❛s ♠♦t✐✈❛t❡❞ ♠❛✐♥❧② ❜② ✐♠♣♦rt❛♥t ❛❞✈❛♥❝❡s ✐♥ ❈♦♠♣✉t❛t✐♦♥❛❧ ❇✐♦❧♦❣② ❛♥❞ t❤❡ ❝♦♥s✐❞❡r❛❜❧❡ ✐♥❝r❡❛s❡ ♦❢ ❞❛t❛❜❛s❡s ♦❢ ❜✐♦❧♦❣✐❝❛❧ ✐♠❛❣❡✳ ■♥ t❤✐s ✇♦r❦ ✐t ✐s ♣r❡s❡♥t❡❞ ❛ ❝♦♠♣✉t❛t✐♦♥❛❧ ♠❡t❤♦❞ t❤❛t ✉s❡s ❞✐✛❡r❡♥t t❡❝❤♥✐q✉❡s ♦❢ ✐♠❛❣❡ ♣r♦❝❡ss✐♥❣✳ ❚❤❡ ♠❛✐♥ ♦❜❥❡❝t✐✈❡ ✐s t♦ ❛❝❤✐❡✈❡ t❤❡ s❡❣♠❡♥t❛t✐♦♥ ♦❢ ❝❡❧❧ ♥✉❝❧❡✐ ❝♦♥t❛✐♥❡❞ ✐♥ s❛❣✐tt❛❧ ✐♠❛❣❡s ♦❢ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r ❡♠❜r②♦s✳ ❚❤✐s t❛s❦ r❡♣r❡s❡♥ts ❛♥ ❡ss❡♥t✐❛❧ st❡♣ ❢♦r ♣❡r❢♦r♠✐♥❣ s❡✈❡r❛❧ ❛♥❛❧②③❡s ❛❜♦✉t t❤✐s ❜✐♦❧♦❣✐❝❛❧ ♠♦❞❡❧✳ ❚❤❡ ♣r♦♣♦s❡❞ ♠❡t❤♦❞ ✐s ❜❛s❡❞ ♦♥ t❤❡ ❡st✐♠❛t✐♦♥ ♦❢ ❝✉r✈❛t✉r❡ ♦❢ t❤❡ ♦❜❥❡❝t ❝♦♥t♦✉r✱ ❛♥❞ ❝♦♥s✐sts ✐♥ s❡♣❛r❛t✐♥❣ ❝❧✉st❡rs ♦❢ ♥✉❝❧❡✐ t❤r♦✉❣❤ s❡❣♠❡♥ts ❞r❛✇♥ ❜❡t✇❡❡♥ ❝♦rr❡s♣♦♥❞✐♥❣ ❝♦♥❝❛✈❡ ♣♦✐♥ts✳ ❚❤✐s ❛♣♣r♦❛❝❤ ✐s ♥❡✇ ❢♦r t❤✐s s❝❡♥❛r✐♦✱ ❛♥❞ ❝♦♥tr✐❜✉t❡s t♦ t❤❡ s♦❧✉t✐♦♥ ♦❢ t❤❡ ♣r♦❜❧❡♠ ♦❢ ❛❣❣r❡❣❛t❡❞ ❝♦♠♣♦♥❡♥ts✱ r❡❝✉rr❡♥t ❢❡❛t✉r❡ ✐♥ ❛♣♣❧✐❝❛t✐♦♥s r❡❧❛t❡❞ t♦ t❤❡ s❡❣♠❡♥t❛t✐♦♥ ♦❢ ♥✉❝❧❡✐✳ ❊✈❛❧✉❛t✐♦♥ ♠❡t❤♦❞s ✐♥❝❧✉❞❡ ❛♣♣r♦❛❝❤❡s t♦ q✉❛♥t✐❢② ❛♥❞ ✈✐s✉❛❧✐③❡ t❤❡ ❛✈❡r❛❣❡ ✐♥t❡♥s✐t② ♦❢ ♣r♦t❡✐♥ ❛♥❞ ❘◆❆ ✐♥ ✐♥tr❛♥✉❝❧❡❛r ❛♥❞ ❝②t♦♣❧❛s♠✐❝ r❡❣✐♦♥s ♦❢ t❤❡ ❡♠❜r②♦ ❉r♦s♦♣❤✐❧❛✳ ❚❤❡ q✉❛♥t✐✜❝❛t✐♦♥ ♠❡t❤♦❞✱ ✐♥ ♣❛rt✐❝✉❧❛r✱ ❛❧❧♦✇s t♦ ♦❜t❛✐♥ t❤❡ ❣❡♥❡ ❡①♣r❡ss✐♦♥ ♣r♦✜❧❡ t❛❦❡♥ ❛❧♦♥❣ t❤❡ ❛♥t❡r✐♦r✲♣♦st❡r✐♦r ❛①✐s ♦❢ t❤❡ ❡♠❜r②♦ ❉r♦s♦♣❤✐❧❛✳ ❖♥ t❤❡ ♦t❤❡r ❤❛♥❞✱ t❤❡ ✈✐s✉❛❧✐③❛t✐♦♥ ♠❡t❤♦❞ ✐♥❝❧✉❞❡s ❛ ♥❡✇ ❛♣♣r♦❛❝❤ ❢♦r t❤❡ ❡①❤✐❜✐t✐♦♥ ♦❢ s❛❣✐tt❛❧ ✐♠❛❣❡s✱ ✐♥ ✇❤✐❝❤ t❤❡ ❝❡❧❧ ♥✉❝❧❡✐ ❛r❡ ❛rr❛♥❣❡❞ ✐♥ ❛ ❧✐♥❡❛r ❝♦♥✜❣✉r❛t✐♦♥✳ ❚❤❡ ❡✣❝✐❡♥t ✉s❡ ♦❢ ❡✈❛❧✉❛t✐♦♥ ♠❡t❤♦❞s ❤❡❧♣s t❤❡ ✐♥t❡r♣r❡t❛t✐♦♥ ♦❢ t❤❡ ❞❛t❛✳ ■♥ ❣❡♥❡r❛❧✱ ♠❡t❤♦❞s ❢♦r ♥✉❝❧❡❛r s❡❣♠❡♥t❛t✐♦♥ ❛♥❞ ❡✈❛❧✉❛t✐♦♥ ♦❢ ❣❡♥❡ ❡①♣r❡ss✐♦♥ r❡♣r❡s❡♥ts ❛♥ ❡ss❡♥t✐❛❧ st❡♣ ❢♦r ♣❡r❢♦r♠✐♥❣ ✈❛r✐♦✉s ❛♥❛❧②③❡s ❜✐♦❧♦❣✐❝❛❧ ♦♥ t❤❡ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ❊①♣❡r✐♠❡♥ts ✇❡r❡ ♣❡r❢♦r♠❡❞ t♦ ✐❧❧✉str❛t❡ t❤❡ ♣❡r❢♦r♠❛♥❝❡ ♦❢ t❤❡ s❡❣♠❡♥t❛t✐♦♥✱ ✇❤❡♥ ❝♦♠♣❛r❡❞ t♦ ♦t❤❡r ❛♣♣r♦❛❝❤❡s ❢♦✉♥❞ ✐♥ t❤❡ ❧✐t❡r❛t✉r❡✳ ❚❤❡ r❡s✉❧ts s❤♦✇❡❞ t❤❛t t❤❡ ♣r♦♣♦s❡❞ ♠❡t❤♦❞ ✐s ❡✛❡❝t✐✈❡ q✉❛❧✐t❛t✐✈❡❧② ❛♥❞ q✉❛♥t✐t❛t✐✈❡❧② ❧❡✈❡❧✳ ❚❤❡ ♠❡t❤♦❞ ❝❛♥ ❛❧s♦ ❜❡ ✉s❡❞ ✐♥ ✈❛r✐♦✉s ❛♣♣❧✐❝❛t✐♦♥s ❛♥❞ ✜❡❧❞s✱ s✉❝❤ t❤❛t ✐♥✈♦❧✈❡ s✐♠✐❧❛r ❝♦♥✜❣✉r❛t✐♦♥s✳

(18)
(19)

❙✉♠ár✐♦

▲✐st❛ ❞❡ ❋✐❣✉r❛s ①①✐

▲✐st❛ ❞❡ ❚❛❜❡❧❛s ①①✈

▲✐st❛ ❞❡ ❆❜r❡✈✐❛t✉r❛s ①①✈✐✐

✶ ■♥tr♦❞✉çã♦ ✷✾

✶✳✶ ❖❜❥❡t✐✈♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✷ ✶✳✶✳✶ ❖❜❥❡t✐✈♦s ❊s♣❡❝í✜❝♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✷ ✶✳✷ Pr✐♥❝✐♣❛✐s ❈♦♥tr✐❜✉✐çõ❡s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✷ ✶✳✸ ❖r❣❛♥✐③❛çã♦ ❞❛ ❞✐ss❡rt❛çã♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✸

✷ ❈♦♥❝❡✐t♦s ❜✐♦❧ó❣✐❝♦s ✸✺

✷✳✶ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✺ ✷✳✷ ❊①♣r❡ssã♦ ●ê♥✐❝❛ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✼ ✷✳✸ ❈❛♣t✉r❛ ❞♦s ❞❛❞♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✸

✸ ❈♦♥❝❡✐t♦s ❈♦♠♣✉t❛❝✐♦♥❛✐s ✹✺

✸✳✶ ❋✉♥❞❛♠❡♥t♦s ❞❡ ■♠❛❣❡♠ ❉✐❣✐t❛❧ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✺ ✸✳✶✳✶ ■♠❛❣❡♠ ❉✐❣✐t❛❧ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✻ ✸✳✶✳✷ ❘❡❧❛❝✐♦♥❛♠❡♥t♦ ❜ás✐❝♦ ❡♥tr❡ ♣✐①❡❧s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✼ ✸✳✶✳✸ ❘♦t✉❧❛çã♦ ❞❡ ❝♦♠♣♦♥❡♥t❡s ❝♦♥❡①♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✾ ✸✳✶✳✹ ❈♦♥t♦r♥♦ ♣❛r❛♠étr✐❝♦ ❞❡ ♦❜❥❡t♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✶ ✸✳✶✳✺ ❇r✐❧❤♦ ❡ ❈♦♥tr❛st❡ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✸ ✸✳✷ ❚é❝♥✐❝❛s ❞❡ ♣ré✲♣r♦❝❡ss❛♠❡♥t♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✸ ✸✳✷✳✶ ❚é❝♥✐❝❛s ❞❡ ❘❡❛❧❝❡ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✹ ✸✳✷✳✷ ❘❡st❛✉r❛çã♦ ❞❡ ✐♠❛❣❡♥s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✺ ✸✳✸ ❚é❝♥✐❝❛s ❞❡ ❙❡❣♠❡♥t❛çã♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✼ ✸✳✸✳✶ ■♥tr♦❞✉çã♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✼ ✸✳✸✳✷ ❙❡❣♠❡♥t❛çã♦ ♣♦r ❙✐♠✐❧❛r✐❞❛❞❡s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✽ ✸✳✸✳✸ ❙❡❣♠❡♥t❛çã♦ ♣♦r ❉❡s❝♦♥t✐♥✉✐❞❛❞❡s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✻✸ ✸✳✹ ▼♦r❢♦❧♦❣✐❛ ▼❛t❡♠át✐❝❛ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✻✻

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①✈✐✐✐ ❙✉♠ár✐♦

✸✳✹✳✶ ❊❧❡♠❡♥t♦ ❊str✉t✉r❛♥t❡ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✻✼ ✸✳✹✳✷ ❖♣❡r❛❞♦r❡s ♠♦r❢♦❧ó❣✐❝♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✻✽ ✸✳✹✳✸ ❚r❛♥s❢♦r♠❛❞❛s ❚♦♣✲❤❛t ❡ ❇♦tt♦♠✲❤❛t ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✼✻ ✸✳✺ ❈✉r✈❛t✉r❛ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✼✼ ✸✳✻ ❆♥á❧✐s❡ ❞❡ ❈♦♠♣♦♥❡♥t❡s Pr✐♥❝✐♣❛✐s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✽✶ ✸✳✼ ❉✐❛❣r❛♠❛s ❞❡ ❱♦r♦♥♦✐ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✽✷ ✸✳✽ ❚r✐❛♥❣✉❧❛çã♦ ❞❡ ❉❡❧❛✉♥❛② ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✽✹

✹ ❚r❛❜❛❧❤♦s ❝♦rr❡❧❛t♦s ✽✼

✹✳✶ ■♥tr♦❞✉çã♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✽✼ ✹✳✷ ❚r❛❜❛❧❤♦s ❝♦rr❡❧❛t♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✽✾ ✹✳✷✳✶ ❙✉♣❡r❢í❝✐❡ ❞♦s ❡♠❜r✐õ❡s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✾✵ ✹✳✷✳✷ P❧❛♥♦ ❛♥❛tô♠✐❝♦ s❛❣✐t❛❧ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✾✶ ✹✳✷✳✸ ❇❛s❡s ❞❡ ❞❛❞♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✾✸ ✺ ■❞❡♥t✐✜❝❛çã♦ ❞❡ ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s ❡♠ ✐♠❛❣❡♥s s❛❣✐t❛✐s ❞❡ ❡♠❜r✐õ❡s ❞❡

❉r♦s♦♣❤✐❧❛ ✾✼

✺✳✶ ▼ét♦❞♦ ♣r♦♣♦st♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✾✼ ✺✳✷ ❆q✉✐s✐çã♦ ❞❛s ✐♠❛❣❡♥s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✾✾ ✺✳✸ Pré✲♣r♦❝❡ss❛♠❡♥t♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✵ ✺✳✸✳✶ ■s♦❧❛♠❡♥t♦ ❞♦ ❡♠❜r✐ã♦ ♣r✐♥❝✐♣❛❧ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✶ ✺✳✸✳✷ ❋✐❧tr❛❣❡♠ ❡s♣❛❝✐❛❧ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✷ ✺✳✸✳✸ ❘❡❛❧❝❡ ❞❡ ❝♦♥tr❛st❡ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✷ ✺✳✸✳✹ ❋✐❧tr❛❣❡♠ ♠♦r❢♦❧ó❣✐❝❛ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✸ ✺✳✹ ❙❡❣♠❡♥t❛çã♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✸ ✺✳✹✳✶ ❈♦♥t♦r♥♦ ♣❛r❛♠étr✐❝♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✼ ✺✳✹✳✷ ❈á❧❝✉❧♦ ❞❛ ❝✉r✈❛t✉r❛ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✽ ✺✳✹✳✸ ❙✉❛✈✐③❛çã♦ ❞❛ ❝✉r✈❛t✉r❛ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✽ ✺✳✹✳✹ ▼í♥✐♠♦s ❧♦❝❛✐s ❞❛ ❝✉r✈❛t✉r❛ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✾ ✺✳✹✳✺ ❈♦rr❡s♣♦♥❞ê♥❝✐❛ ❡♥tr❡ ♦s ♣♦♥t♦s ❞❡ ♠í♥✐♠♦s ❧♦❝❛✐s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✵✾ ✺✳✹✳✻ ❙❡♣❛r❛çã♦ ❞♦s ♥ú❝❧❡♦s ❛❣r❡❣❛❞♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✶✷ ✻ ◗✉❛♥t✐✜❝❛çã♦ ❡ ❱✐s✉❛❧✐③❛çã♦ ❞❛ ❊①♣r❡ssã♦ ●ê♥✐❝❛ ❡♠ ✐♠❛❣❡♥s s❛❣✐t❛✐s

❞❡ ❉r♦s♦♣❤✐❧❛ ✶✶✺

✻✳✶ ◆í✈❡❧ ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ♣♦r ♥ú❝❧❡♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✶✺ ✻✳✷ ◆í✈❡❧ ❞❡ ❡①♣r❡ssã♦ ♥❛ r❡❣✐ã♦ ❞♦ ❝✐t♦♣❧❛s♠❛ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✶✻ ✻✳✸ ❱✐s✉❛❧✐③❛çã♦ ❞❛ ✈❛r✐❛çã♦ ❞❡ ✐♥t❡♥s✐❞❛❞❡ ❞❡ ❡①♣r❡ssã♦ ♥✉❝❧❡❛r ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✶✽

✼ ❘❡s✉❧t❛❞♦s ❡ ❉✐s❝✉ssã♦ ✶✷✺

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❙✉♠ár✐♦ ①✐①

✼✳✶✳✶ ❚r❛t❛♠❡♥t♦ ❞❛s ✐♠❛❣❡♥s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✷✺ ✼✳✷ ❊t❛♣❛ ❞❡ s❡❣♠❡♥t❛çã♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✸✶ ✼✳✷✳✶ ❘❡s✉❧t❛❞♦s ❞❛ ❜✐♥❛r✐③❛çã♦ ❞❛ ✐♠❛❣❡♠ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✸✶ ✼✳✷✳✷ ❘❡s✉❧t❛❞♦s ❞♦ ♣ós✲♣r♦❝❡ss❛♠❡♥t♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✸✺ ✼✳✸ ❆✈❛❧✐❛çã♦ ❡ ❛♥á❧✐s❡ ❝♦♠♣❛r❛t✐✈❛ ❞♦s r❡s✉❧t❛❞♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✵ ✼✳✸✳✶ ❇❛s❡s ❞❡ ❞❛❞♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✵ ✼✳✸✳✷ ▼ét♦❞♦s ❛✈❛❧✐❛❞♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✵ ✼✳✸✳✸ ▼❡❞✐❞❛s ❞❡ ❛✈❛❧✐❛çã♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✶ ✼✳✹ ❊t❛♣❛ ❞❡ ◗✉❛♥t✐✜❝❛çã♦ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✺✶

✽ ❈♦♥❝❧✉sã♦ ✶✻✶

✽✳✶ ❚r❛❜❛❧❤♦s ❋✉t✉r♦s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✻✷ ✽✳✷ P✉❜❧✐❝❛çõ❡s r❡❛❧✐③❛❞❛s ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✻✸

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(23)

▲✐st❛ ❞❡ ❋✐❣✉r❛s

✶✳✶ ▼ás❝❛r❛ ❞❡ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✶ ✶✳✷ ❊♠❜r✐ã♦ ❞❡ ❉r♦s♦♣❤✐❧❛ ♥♦ ♣❧❛♥♦ s❛❣✐t❛❧✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✷ ✷✳✶ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r ❢ê♠❡❛✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✻ ✷✳✷ ❈✐❝❧♦s ❞❡ ❝❧✐✈❛❣❡♠ ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✼ ✷✳✸ ❉✐✈❡rs✐❞❛❞❡ ♠♦r❢♦❧ó❣✐❝❛ ❞❡ ❝é❧✉❧❛s✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✸✾ ✷✳✹ P❛❞rã♦ ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✶ ✷✳✺ ❱✐s✉❛❧✐③❛çã♦ ❣rá✜❝❛ ❞♦s ♣❧❛♥♦s ❛♥❛tô♠✐❝♦s ❞❛ ❉r♦s♦♣❤✐❧❛✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✷ ✷✳✻ ■♠❛❣❡♥s s❛❣✐t❛✐s ❞❡ ❡♠❜r✐õ❡s ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✷ ✸✳✶ P❛ss♦s ❢✉♥❞❛♠❡♥t❛✐s ❡♠ ♣r♦❝❡ss❛♠❡♥t♦ ❞✐❣✐t❛❧ ❞❡ ✐♠❛❣❡♥s✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✻ ✸✳✷ ■❧✉str❛çã♦ ❞♦s ♣r♦❝❡ss♦s ❞❡ ❛♠♦str❛❣❡♠ ❡ q✉❛♥t✐③❛çã♦✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✽ ✸✳✸ ❚✐♣♦s ❞❡ ✈✐③✐♥❤❛♥ç❛✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✽ ✸✳✹ ❚✐♣♦s ❞❡ ❝♦♥❡❝t✐✈✐❞❛❞❡✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✹✾ ✸✳✺ ❘♦t✉❧❛çã♦ ❞❡ ♦❜❥❡t♦s ❝♦♥❡①♦s✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✵ ✸✳✻ ❆❧❣♦r✐t♠♦ ❙❡❣✉✐♠❡♥t♦ ❞❡ ❝♦♥t♦r♥♦ ✭❈❡❣✉✐♥❤♦✮✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✷ ✸✳✼ ❱✐③✐♥❤❛♥ç❛ ✽✲❝♦♥❡❝t❛❞♦ ❞❡p ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✷

✸✳✽ ▼ás❝❛r❛ ❞❡ ❝♦♥✈♦❧✉çã♦ ❣❛✉ss✐❛♥❛✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✺✼ ✸✳✾ ❊❧❡♠❡♥t♦ ❡str✉t✉r❛♥t❡ ♥♦ ❢♦r♠❛t♦ ✏❝r✉③✑✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✻✼ ✸✳✶✵ ❊❧❡♠❡♥t♦ ❡str✉t✉r❛♥t❡ ♥♦ ❢♦r♠❛t♦ ✏❞✐❛♠❛♥t❡✑✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✻✽ ✸✳✶✶ ❊①❡♠♣❧♦ ❞❡ ❡r♦sã♦ ❜✐♥ár✐❛✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✼✶ ✸✳✶✷ ❊①❡♠♣❧♦ ❞❡ ❞✐❧❛t❛çã♦ ❜✐♥ár✐❛✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✼✷ ✸✳✶✸ ❖ ❢❛♠♦s♦ ✏❆tt♥❡❛✈❡✬s ❝❛t✑✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✼✽ ✸✳✶✹ ❘❡♣r❡s❡♥t❛çã♦ ❞♦ ❝♦♥t♦r♥♦ ❞❡ ✉♠❛ ❢♦r♠❛ ❝♦♠ ❞✐❢❡r❡♥t❡s ✈❛❧♦r❡s ❞❡ σ✳ ✳ ✳ ✳ ✽✵

✸✳✶✺ ❊♥t✐❞❛❞❡s t♦♣♦❧ó❣✐❝❛s ❞♦ ❉✐❛❣r❛♠❛ ❞❡ ❱♦r♦♥♦✐✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✽✸ ✸✳✶✻ ❚r✐❛♥❣✉❧❛çã♦ ❞❡ ❉❡❧❛✉♥❛② ❝♦♠ ❝ír❝✉❧♦s✲❝✐r❝✉♥❞❛♥t❡s✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✽✹ ✸✳✶✼ ❉✐❛❣r❛♠❛ ❞❡ ❱♦r♦♥♦✐ ❡ ❚r✐❛♥❣✉❧❛çã♦ ❞❡ ❉❡❧❛✉♥❛②✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✽✺ ✹✳✶ ■❞❡♥t✐✜❝❛çã♦ ❞❡ ♥ú❝❧❡♦s ❝♦♠ ❜❛s❡ ❡♠ ❥❛♥❡❧❛ ❞❡s❧✐③❛♥t❡ r❡t❛♥❣✉❧❛r✳ ✳ ✳ ✳ ✳ ✳ ✾✷ ✹✳✷ ■❞❡♥t✐✜❝❛çã♦ ❞❡ ♥ú❝❧❡♦s ❝♦♠ ❜❛s❡ ❡♠ ❥❛♥❡❧❛ ❞❡s❧✐③❛♥t❡ ❝✐r❝✉❧❛r✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✾✷ ✺✳✶ ❊sq✉❡♠❛ ❣❡r❛❧ ❞❛ ♠❡t♦❞♦❧♦❣✐❛ ♣r♦♣♦st❛✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✾✽

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①①✐✐ ▲✐st❛ ❞❡ ❋✐❣✉r❛s

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▲✐st❛ ❞❡ ❋✐❣✉r❛s ①①✐✐✐

✼✳✶✻ ❘❡s✉❧t❛❞♦s ♦❜t✐❞♦s ♣❡❧♦ ♠ét♦❞♦ ♣r♦♣♦st♦✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✻ ✼✳✶✼ ❘❡s✉❧t❛❞♦s ♦❜t✐❞♦s ♣❡❧♦ ♠ét♦❞♦ ♣r♦♣♦st♦✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✼ ✼✳✶✽ ❘❡s✉❧t❛❞♦s ♦❜t✐❞♦s ♣❡❧♦ ♠ét♦❞♦ ♣r♦♣♦st♦✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✽ ✼✳✶✾ ❘❡s✉❧t❛❞♦s ♦❜t✐❞♦s ♣❡❧♦ ♠ét♦❞♦ ♣r♦♣♦st♦✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✾ ✼✳✷✵ ❘❡s✉❧t❛❞♦s ♦❜t✐❞♦s ♣❡❧♦ ♠ét♦❞♦ ♣r♦♣♦st♦✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✺✵ ✼✳✷✶ ❙❡♣❛r❛çã♦ ❞❛s r❡❣✐õ❡s ♥✉❝❧❡❛r ❡ ❝✐t♦♣❧❛s♠❛✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✺✶ ✼✳✷✷ ❊♠❜r✐ã♦ ✶ ✲ P❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❡ ✐♥t❡♥s✐❞❛❞❡s ❞❡ ✢✉♦r❡s❝ê♥❝✐❛ ♥♦s ♥ú❝❧❡♦s✳ ✶✺✷ ✼✳✷✸ ❊♠❜r✐ã♦ ✷ ✲ P❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❡ ✐♥t❡♥s✐❞❛❞❡s ❞❡ ✢✉♦r❡s❝ê♥❝✐❛ ♥♦s ♥ú❝❧❡♦s✳ ✶✺✸ ✼✳✷✹ ❊♠❜r✐ã♦ ✸ ✲ P❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❡ ✐♥t❡♥s✐❞❛❞❡s ❞❡ ✢✉♦r❡s❝ê♥❝✐❛ ♥♦s ♥ú❝❧❡♦s✳ ✶✺✹ ✼✳✷✺ ●rá✜❝♦ ❝♦♠♣❛r❛t✐✈♦ ❞♦ ♣❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❞❛ ♣r♦t❡í♥❛ ❍✉♥❝❤❜❛❝❦✳ ✳ ✳ ✳ ✳ ✳ ✶✺✺ ✼✳✷✻ ●rá✜❝♦ ❞❡ ▼é❞✐❛s ♣❛r❛ ♦ ❝♦♥❥✉♥t♦ ❞❡ ✐♠❛❣❡♥s ❞❛ ♣r♦t❡í♥❛ ❍✉♥❝❤❜❛❝❦✳ ✳ ✳ ✶✺✺ ✼✳✷✼ ❊♠❜r✐ã♦ ✶ ✲ P❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❡ ✐♥t❡♥s✐❞❛❞❡s ❞❡ ✢✉♦r❡s❝ê♥❝✐❛ ❞♦ ❝✐t♦♣❧❛s♠❛✳✶✺✻ ✼✳✷✽ ❊♠❜r✐ã♦ ✷ ✲ P❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❡ ✐♥t❡♥s✐❞❛❞❡s ❞❡ ✢✉♦r❡s❝ê♥❝✐❛ ❞♦ ❝✐t♦♣❧❛s♠❛✳✶✺✼ ✼✳✷✾ ❊♠❜r✐ã♦ ✸ ✲ P❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❡ ✐♥t❡♥s✐❞❛❞❡s ❞❡ ✢✉♦r❡s❝ê♥❝✐❛ ❞♦ ❝✐t♦♣❧❛s♠❛✳✶✺✽ ✼✳✸✵ ●rá✜❝♦ ❝♦♠♣❛r❛t✐✈♦ ❞♦ ♣❡r✜❧ ❞❡ ❡①♣r❡ssã♦ ❞♦ ❘◆❆ ❞❡ ❉r♦s♦♣❤✐❧❛✳ ✳ ✳ ✳ ✳ ✳ ✶✺✾ ✼✳✸✶ ●rá✜❝♦ ❞❡ ▼é❞✐❛s ♣❛r❛ ♦ ❝♦♥❥✉♥t♦ ❞❡ ✐♠❛❣❡♥s ❝♦♠ s✐♥❛❧ ❞♦ ❘◆❆ ❞❡ ❍✉♥✲

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▲✐st❛ ❞❡ ❚❛❜❡❧❛s

✼✳✶ ❘❡s✉❧t❛❞♦s ❞❛ ❛♣❧✐❝❛çã♦ ❞♦ ♠ét♦❞♦ ♣r♦♣♦st♦✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✸ ✼✳✷ ❈♦♠♣❛r❛çã♦ ❞♦s r❡s✉❧t❛❞♦s ♣❛r❛ ❞✐❢❡r❡♥t❡s ♠ét♦❞♦s✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✳ ✶✹✹

(28)
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▲✐st❛ ❞❡ ❆❜r❡✈✐❛t✉r❛s

❆P ❆♥t❡r✐♦r✲P♦st❡r✐♦r✳ ✷❉ ❇✐❞✐♠❡♥s✐♦♥❛❧✳ ✸❉ ❚r✐❞✐♠❡♥s✐♦♥❛❧✳

❇❉●P ❇❡r❦❡❧❡② ❉r♦s♦♣❤✐❧❛ ●❡♥♦♠❡ Pr♦❥❡❝t

❇❉❚◆P ❇❡r❦❡❧❡② ❉r♦s♦♣❤✐❧❛ ❚r❛♥s❝r✐♣t✐♦♥ ◆❡t✇♦r❦ Pr♦❥❡❝t ❉◆❆ ➪❝✐❞♦ ❞❡s♦①✐rr✐❜♦♥✉❝❧❡✐❝♦ ✭❉❡s♦①②r✐❜♦♥✉❝❧❡✐❝ ❛❝✐❞✮✳ ❊❙ ❊❧❡♠❡♥t♦ ❡str✉t✉r❛♥t❡✳

❊❙s ❊❧❡♠❡♥t♦s ❡str✉t✉r❛♥t❡s✳

❊q✳ ❊q✉❛çã♦✳

❊qs✳ ❊q✉❛çõ❡s✳ ❋✐❣✳ ❋✐❣✉r❛✳ ❋✐❣s✳ ❋✐❣✉r❛s✳

❋❚ ❚r❛♥s❢♦r♠❛❞❛ ❞❡ ❋♦✉r✐❡r ✭❋♦✉r✐❡r ❚r❛♥s❢♦r♠✮✳ ▼❈ ▼✐❝r♦s❝♦♣✐❛ ❈♦♥❢♦❝❛❧✳

▼▼ ▼♦r❢♦❧♦❣✐❛ ▼❛t❡♠át✐❝❛

P❈❆ ❆♥á❧✐s❡ ❞❡ ❝♦♠♣♦♥❡♥t❡s ♣r✐♥❝✐♣❛✐s ✭Pr✐♥❝✐♣❛❧ ❈♦♠♣♦♥❡♥t ❆♥❛❧②s✐s✮✳ P❉■ Pr♦❝❡ss❛♠❡♥t♦ ❉✐❣✐t❛❧ ❞❡ ■♠❛❣❡♥s

❘◆❆ ➪❝✐❞♦ r✐❜♦♥✉❝❧❡✐❝♦ ✭❘✐❜♦♥✉❝❧❡✐❝ ❛❝✐❞✮✳ ❙❡❝✳ ❙❡çã♦✳

❚❉ ❚r❛♥s❢♦r♠❛❞❛ ❉✐stâ♥❝✐❛

❚■❋❋ ❋♦r♠❛t♦ ❞❡ ❛rq✉✐✈♦ ❞❡ ✐♠❛❣❡♠ ♠❛r❝❛❞♦ ✭❚❛❣❣❡❞ ■♠❛❣❡ ❋✐❧❡ ❋♦r♠❛t✮✳

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❈❛♣ít✉❧♦ ✶

■♥tr♦❞✉çã♦

◆♦s ú❧t✐♠♦s ❛♥♦s✱ ❛ ❇✐♦✐♥❢♦r♠át✐❝❛ ❡ ❛ ❇✐♦❧♦❣✐❛ ❈♦♠♣✉t❛❝✐♦♥❛❧ t♦r♥❛r❛♠✲s❡ ❡ss❡♥❝✐❛✐s ♣❛r❛ ❛ r❡❛❧✐③❛çã♦ ❞❡ ❡st✉❞♦s ❜✐♦❧ó❣✐❝♦s✱ s❡♥❞♦ ❝r❡s❝❡♥t❡ ♦ ❞❡s❡♥✈♦❧✈✐♠❡♥t♦ ❞❡ ❢❡rr❛♠❡♥t❛s ❡ ♠ét♦❞♦s ❝♦♠♣✉t❛❝✐♦♥❛✐s ❝❛♣❛③❡s ❞❡ ♣r♦♠♦✈❡r ❛ ❛q✉✐s✐çã♦✱ ♣r♦❝❡ss❛♠❡♥t♦✱ ❛r♠❛③❡♥❛✲ ♠❡♥t♦✱ ❞✐str✐❜✉✐çã♦✱ ❛♥á❧✐s❡ ❡ ✐♥t❡r♣r❡t❛çã♦ ❞❛ ✐♥❢♦r♠❛çã♦ ❜✐♦❧ó❣✐❝❛✳ P❛r❛ ❬❍❛❣❡♥ ✷✵✵✵❪✱ ❡ss❡ ❝r❡s❝✐♠❡♥t♦ ❢♦✐ ♠♦t✐✈❛❞♦✱ ♣r✐♥❝✐♣❛❧♠❡♥t❡✱ ♣♦r ❛✈❛♥ç♦s ✐♠♣♦rt❛♥t❡s ♥♦s ♠ét♦❞♦s ❞❡ s❡q✉❡♥❝✐❛♠❡♥t♦ ❞❡ ❉◆❆ ✭á❝✐❞♦ ❞❡s♦①✐rr✐❜♦♥✉❝❧❡✐❝♦✮✱ s✉r❣✐♠❡♥t♦ ❞♦s ♣r♦❥❡t♦s ❣❡♥♦♠❛ ✭q✉❡ ❣❡r❛r❛♠ ✈♦❧✉♠❡s ✐♥t❡♥s♦s ❞❡ ✐♥❢♦r♠❛çõ❡s✮✱ tr❛♥s❝r✐♣t♦♠❛ ❡ ♣r♦t❡♦♠❛✱ ❞❡s❡♥✈♦❧✈✐✲ ♠❡♥t♦ ❞❡ ♠✐❝r♦❛rr❛♥❥♦s ❡ ❜✐♦❝❤✐♣s✱ ❡ ♣❡❧♦ ❛✉♠❡♥t♦ ❞❡ ❜❛♥❝♦s ❞❡ ❞❛❞♦s ❜✐♦❧ó❣✐❝♦s✳ ❆❧✐❛❞♦ ❛ ❡ss❡s ❢❛t♦r❡s✱ ❝✐t❛♠✲s❡ ❛ ❞✐s♣♦♥✐❜✐❧✐❞❛❞❡ ❞❡ ❝♦♠♣✉t❛❞♦r❡s ♠❛✐s ✈❡❧♦③❡s ❡ r♦❜✉st♦s✱ ❛❧é♠ ❞♦ ❛✈❛♥ç♦ ❞❛ ■♥t❡r♥❡t✳

❊♠ ♣❛rt✐❝✉❧❛r✱ ❛ ❛♣❧✐❝❛çã♦ ❞❡ té❝♥✐❝❛s ❝♦♠♣✉t❛❝✐♦♥❛✐s✱ ❡s♣❡❝✐❛❧♠❡♥t❡ ❡♠ ❡st✉❞♦s ❞❛ ❇✐♦❧♦❣✐❛ ▼♦❧❡❝✉❧❛r✱ ♣♦ss✐❜✐❧✐t❛ ♦ ❡st✉❞♦ ❞❛ ❡str✉t✉r❛✱ ♦r❣❛♥✐③❛çã♦ ❡ ❡✈♦❧✉çã♦ ❞❡ ❣❡♥♦✲ ♠❛s❀ ❡st✉❞♦ ❞❛ ❡①♣r❡ssã♦ ❞✐❢❡r❡♥❝✐❛❧ ❞❡ ❣❡♥❡s ❡ ♣r♦t❡í♥❛s❀ ❛ ♣r❡❞✐çã♦ ❞❛ ❝♦♥✜❣✉r❛çã♦ tr✐❞✐♠❡♥s✐♦♥❛❧ ❞❡ ♣r♦t❡í♥❛s❀ ❛♥á❧✐s❡ ❞❛ ❡str✉t✉r❛✱ ❢✉♥çã♦ ❡ ❡✈♦❧✉çã♦ ♠♦❧❡❝✉❧❛r❡s✱ ❡♥tr❡ ♦✉tr♦s ❬❈❛tt❧❡② ❛♥❞ ❆rt❤✉r ✷✵✵✼❪✳ ❙♦❜ ❡ss❡ ❛s♣❡❝t♦✱ ♦ ❞❡s❡♥✈♦❧✈✐♠❡♥t♦ ❞❡ ♠ét♦❞♦s ❝♦♠✲ ♣✉t❛❝✐♦♥❛✐s t❡♠ ♣❡r♠✐t✐❞♦ ❛♦s ❜✐♦❧♦❣✐st❛s ♠♦❧❡❝✉❧❛r❡s ♣r♦❝❡ss❛r✱ ✐♥t❡r♣r❡t❛r ❡ ❛♥❛❧✐s❛r ❛✉t♦♠❛t✐❝❛♠❡♥t❡ ❣r❛♥❞❡s q✉❛♥t✐❞❛❞❡s ❞❡ ❞❛❞♦s ❜✐♦❧ó❣✐❝♦s✳ ❙❡❣✉♥❞♦ ❬❖✉③♦✉♥✐s ✷✵✵✷❪ ❞❡st❛❝❛♠✲s❡ ♦s ❛❧❣♦r✐t♠♦s ♣❛r❛ ❛❧✐♥❤❛♠❡♥t♦ ❞❡ s❡q✉ê♥❝✐❛s✱ ♣❛r❛ ❛ ❝r✐❛çã♦ ❞❡ ❜❛♥❝♦s ❞❡ ❞❛❞♦s ❞❡ s❡q✉ê♥❝✐❛s ♣r♦t❡✐❝❛s ✭❞❡ ❛❝❡ss♦ ♣ú❜❧✐❝♦ ♦✉ ♣r✐✈❛❞♦✮✱ ♣❛r❛ ❛♥á❧✐s❡ ❞❡ ♣❛❞rõ❡s ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛✱ ❡ ♣❛r❛ ❛♥♦t❛çã♦ ❡ ❝♦♠♣❛r❛çã♦ ❞❡ ❣❡♥♦♠❛s✳

◆❡st❡ ❝❡♥ár✐♦✱ ♦s ❛❧❣♦r✐t♠♦s ✈♦❧t❛❞♦s à ✐❞❡♥t✐✜❝❛çã♦ ❡ ❛♥á❧✐s❡ ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ t❡♠ ❛tr❛í❞♦ ❣r❛♥❞❡ ❛t❡♥çã♦ ❞❛ ❝♦♠✉♥✐❞❛❞❡ ❝✐❡♥tí✜❝❛✱ s❡♥❞♦ ❛♠♣❧❛♠❡♥t❡ ✉t✐❧✐③❛❞♦s ❡♠ ♣❡s✲ q✉✐s❛s ❜✐♦❧ó❣✐❝❛s ❬▲♦r❦♦✇s❦❧ ❛♥❞ ❈✉❧❧❡♥ ✷✵✵✹✱ ◗✉❛❝❦❡♥❜✉s❤ ✷✵✵✶✱ ❞❡ ❆r❛ú❥♦ ✷✵✵✽❪✳ ■ss♦ ♣♦rq✉❡✱ ❛ ❛♥á❧✐s❡ ❝♦♠♣✉t❛❝✐♦♥❛❧ ♣❡r♠✐t❡ ♦❜t❡r✱ ❞❡ ❢♦r♠❛ ❛✉t♦♠át✐❝❛ ❡ ♣r❡❝✐s❛✱ r❡❧❡✈❛♥t❡s ✐♥❢♦r♠❛çõ❡s s♦❜r❡ ♦ ❢✉♥❝✐♦♥❛♠❡♥t♦ ❞❡ ✉♠❛ ❝é❧✉❧❛✱ t❛✐s ❝♦♠♦✱ ❢✉♥çã♦ ❞❡ ❝❛❞❛ ❣❡♥❡ ♦✉ ♣r♦t❡í♥❛ ❝♦❞✐✜❝❛❞❛✱ ❝♦♥tr♦❧❡ ❞❛ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❡ s✉❛s ✐♥t❡r❛çõ❡s s♦❜ ❞❡t❡r♠✐♥❛❞❛s s✐✲ t✉❛çõ❡s✱ ✐❞❡♥t✐✜❝❛çã♦ ❞❡ ❣❡♥❡s ❞✐❢❡r❡♥❝✐❛❧♠❡♥t❡ ❡①♣r❡ss♦s ❡♠ ❝♦♥❞✐çõ❡s ❞✐st✐♥t❛s✱ ❞❡♥tr❡ ♦✉tr❛s ❬❨❛♥❣ ❡t ❛❧✳ ✷✵✵✻❜❪✳

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✸✵ ❈❛♣ít✉❧♦ ✶✳ ■♥tr♦❞✉çã♦

❯♠ ❡①❡♠♣❧♦ ✐♠♣♦rt❛♥t❡✱ é ❛ ❛♥á❧✐s❡ ❞♦s ♣❛❞rõ❡s ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ❊st✉❞♦s ❞❡ ❜✐♦✐♥❢♦r♠át✐❝❛ s♦❜r❡ ❡ss❡ ♠♦❞❡❧♦ ❜✐♦❧ó❣✐❝♦ ❢♦r♥❡❝❡♠ ❛s ❜❛s❡s ♣❛r❛ ❝♦♠♣r❡❡♥sã♦ ❞❡ ♣r♦❝❡ss♦s ❡ ❢❡♥ô♠❡♥♦s ❞❡ ❞✐✈❡rs♦s ♦r❣❛♥✐s♠♦s✱ t❛✐s ❝♦♠♦ ❞❡s❡♥✈♦❧✈✐✲ ♠❡♥t♦ ❡ ❞✐❢❡r❡♥❝✐❛çã♦ ❝❡❧✉❧❛r ❬❙❡♣❡❧ ❛♥❞ ▲♦r❡t♦ ✷✵✶✵✱●✐❧❜❡rt ✷✵✵✸❪✳ ❆❧é♠ ❞✐ss♦✱ ♦ ❣r❛✉ ❞❡ ❝♦♥s❡r✈❛çã♦ ❜✐♦❧ó❣✐❝❛ ❡ ✜s✐♦❧ó❣✐❝❛ ❝♦♠ ♦ ❣❡♥♦♠❛ ❤✉♠❛♥♦ ❢❛✈♦r❡❝❡ ❛ ❝♦♠♣r❡❡♥sã♦ ❞♦s ♠❡✲ ❝❛♥✐s♠♦s ❞❡ tr❛♥s♠✐ssã♦ ❞❡ ❝❛r❛❝t❡ríst✐❝❛s ❤❡r❡❞✐tár✐❛s ❡ ✐❞❡♥t✐✜❝❛çã♦ ❞❡ ❣❡♥❡s ❛ss♦❝✐❛❞♦s ❛ ❞✐stúr❜✐♦s ❞✐✈❡rs♦s ✭❝♦♠♦ ❝â♥❝❡r✱ ♠❛❧ ❞❡ ❆❧③❤❡✐♠❡r✱ ❝♦♠♣♦rt❛♠❡♥t♦s ❞❡ ❛♣r❡♥❞✐③❛❣❡♠✱ P❛r❦✐♥s♦♥ ❡ ❛❧❝♦♦❧✐s♠♦✮ ❬❈❤❡♥ ❛♥❞ ❈r♦✇t❤❡r ✷✵✶✷✱ ❍✐rt❤ ✷✵✶✵❪✳ ❚❛✐s ❢❛t♦r❡s t♦r♥❛r❛♠ ❛ ❉r♦s♦♣❤✐❧❛ ✉♠ ✐♠♣♦rt❛♥t❡ ♠♦❞❡❧♦ ♣❛r❛ ❛ ✐❞❡♥t✐✜❝❛çã♦✱ ❝❛r❛❝t❡r✐③❛çã♦ ❡ ♦ ❡st✉❞♦ ❞♦s ♣❛❞rõ❡s ❞❡ ❞✐str✐❜✉✐çã♦ ❣ê♥✐❝❛✳

❯♠❛ ❞❛s ❢♦r♠❛s ❞❡ r❡❛❧✐③❛r ♦ ♣r♦❝❡ss❛♠❡♥t♦ ❡ ✐♥t❡r♣r❡t❛çã♦ ❞❡ ❞❛❞♦s ❞❛ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❡♥✈♦❧✈❡ ❛ ♠❛♥✐♣✉❧❛çã♦ ❞❡ ✐♠❛❣❡♥s ❞✐❣✐t❛❧✐③❛❞❛s✱ ♣❛r❛ ✜♥s ❞❡ ❛✈❛❧✐❛çã♦ ❡ q✉❛♥t✐✲ ✜❝❛çã♦ ❞❛s ♣r♦t❡í♥❛s ❡ ❘◆❆s ✭á❝✐❞♦ r✐❜♦♥✉❝❧❡✐❝♦✮✳ ❊♥tr❡t❛♥t♦✱ ❡ss❛ ♥ã♦ é ✉♠❛ t❛r❡❢❛ tr✐✈✐❛❧✱ ❥á q✉❡ ❡♥✈♦❧✈❡ ❛ ✐❞❡♥t✐✜❝❛çã♦ ❡ ❡st✉❞♦ s✐♠✉❧tâ♥❡♦ ❞❡ ✉♠ ❣r❛♥❞❡ ♥ú♠❡r♦ ❞❡ ❣❡✲ ♥❡s ❬❈❛❧s❛ ❏✉♥✐♦r ❡t ❛❧✳ ✷✵✵✹❪✳ ❆❧é♠ ❞✐ss♦✱ ♣❛r❛ ❛ ♠❛✐♦r✐❛ ❞❛s ✐♥✈❡st✐❣❛çõ❡s ❜✐♦❧ó❣✐❝❛s✱ ♥♦r♠❛❧♠❡♥t❡ é ♥❡❝❡ssár✐♦ ❝❛r❛❝t❡r✐③❛r ❞✐❢❡r❡♥t❡s ♣❛❞rõ❡s ❡s♣❛ç♦✲t❡♠♣♦r❛✐s ❞❛ ❡①♣r❡ssã♦ ❣ê♥✐❝❛✱ ❡♠ ❞✐❢❡r❡♥t❡s ❡stá❣✐♦s ❞♦ ❞❡s❡♥✈♦❧✈✐♠❡♥t♦ ❡♠❜r✐♦♥ár✐♦✳

❋r❡♥t❡ ❛ ❡ss❛ ❝♦♠♣❧❡①✐❞❛❞❡✱ t♦r♥❛✲s❡ ♥❡❝❡ssár✐♦ ♦ ✉s♦ ❞❡ té❝♥✐❝❛s ❡✴♦✉ ❢❡rr❛♠❡♥t❛s ❝♦♠♣✉t❛❝✐♦♥❛✐s ♣❛r❛ ❛ ♦r❣❛♥✐③❛çã♦ ❡ ♣r♦❝❡ss❛♠❡♥t♦ ❞♦s ❞❛❞♦s✱ ❜❡♠ ❝♦♠♦ ♥♦ ❛✉①í❧✐♦ ❞❡ ♥♦✈❛s ❞❡s❝♦❜❡rt❛s ❬❚r❛✈❡♥ç♦❧♦ ✷✵✵✼❪✳ ◆❡st❡ s❡♥t✐❞♦✱ é ♣♦ssí✈❡❧ ❡♥❝♦♥tr❛r ✈❛r✐❛❞♦s ♠ét♦❞♦s ♣r♦♣♦st♦s ♥❛ ❧✐t❡r❛t✉r❛✳ ❆ ♠❛✐♦r✐❛ s❡ ❞✐❢❡r❡♥❝✐❛ ❝♦♠ r❡❧❛çã♦ ❛♦ t✐♣♦ ❞❡ té❝♥✐❝❛✴❛❧❣♦r✐t♠♦ ✉t✐❧✐③❛❞♦✱ ♦✉ ❝♦♠ r❡❧❛çã♦ ❛♦ t✐♣♦ ❞❡ ✐♥✈❡st✐❣❛çã♦ ❜✐♦❧ó❣✐❝❛ ❞❡ ✐♥t❡r❡ss❡✳ ❖ ❣r❛♥❞❡ ❞❡s❛✜♦ ❡♥❝♦♥tr❛❞♦ ♥❡ss❡s tr❛❜❛❧❤♦s é ❛ t❛r❡❢❛ ❞❡ s❡❣♠❡♥t❛çã♦ ❛✉t♦♠át✐❝❛ ❞♦s ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s ❝♦♥t✐❞♦s ♥❛ ❜❧❛st♦❞❡r♠❡ ❞❡ ❡♠❜r✐õ❡s ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ❖ ♦❜❥❡t✐✈♦ ❞❡st❛ t❛r❡❢❛ é ♦❜t❡r ❛ ♠ás❝❛r❛ ❞❡ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r ✭❝♦♠♦ ❛ ♠♦str❛❞❛ ♥❛ ❋✐❣✳✶✳✷✮✱ ✐st♦ é✱ ✉♠❛ ✐♠❛❣❡♠ ❜✐♥ár✐❛ ❡♠ q✉❡ ♦s ♣✐①❡❧s ❝♦♠ ✈❛❧♦r ✐❣✉❛❧ ❛ ✶ ✭✉♠✮ r❡♣r❡s❡♥t❛♠ ♦s ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s ❞♦ ❡♠❜r✐ã♦✱ ❡ ♦s ♣✐①❡❧s ❝♦♠ ✈❛❧♦r ✐❣✉❛❧ ❛ ✵ ✭③❡r♦✮ r❡♣r❡s❡♥t❛♠ ❛s r❡❣✐õ❡s ❢♦r❛ ❞♦ ♥ú❝❧❡♦ ❡ ♦ ❢✉♥❞♦ ❞❛ ✐♠❛❣❡♠✳

❆ ♠ás❝❛r❛ ❞❡ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r é ♣r✐♠♦r❞✐❛❧ ♣❛r❛ ❛ r❡❛❧✐③❛çã♦ ❞❡ ❞✐✈❡rs❛s t❛r❡❢❛s s✉❜s❡q✉❡♥t❡s✱ t❛✐s ❝♦♠♦✱ q✉❛♥t✐✜❝❛çã♦ ❞♦ ♥í✈❡❧ ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❡♠ ❝❛❞❛ ♥ú❝❧❡♦ ❬P❛♥ ❡t ❛❧✳ ✷✵✵✻❪✱ ❞❡t❡❝çã♦ ❞❛ ✐❞❛❞❡ ❞❡ ❡♠❜r✐õ❡s ❞❡ ❉r♦s♦♣❤✐❧❛ ❬❆✐③❡♥❜❡r❣ ❡t ❛❧✳ ✷✵✵✷❪✱ ❛♥á❧✐s❡ ❞❛s r❡❞❡s ❞❡ r❡❣✉❧❛çã♦ ❣ê♥✐❝❛✱ ❝♦♠♣❛r❛çã♦ ❞♦s ♣❛❞rõ❡s ❞❡ ❡①♣r❡ssã♦ ❡♠ ❞✐❢❡r❡♥t❡s ❡stá❣✐♦s ❞♦ ❞❡s❡♥✈♦❧✈✐♠❡♥t♦ ❬❑✉♠❛r ❡t ❛❧✳ ✷✵✵✷❪✱ ❛♥á❧✐s❡ ❞✐♥â♠✐❝❛ ❞❛ ❢♦r♠❛çã♦ ❞♦s ♣❛❞rõ❡s ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❬P♦✉st❡❧♥✐❦♦✈❛ ❡t ❛❧✳ ✷✵✵✹❪✱ ❛♥á❧✐s❡ ❞♦ ❝♦♠♣♦rt❛♠❡♥t♦ ❞❡ ✉♠ ❣❡♥❡ ❡♠ ❢✉♥çã♦ ❞❡ ♦✉tr♦ ❬❋♦✇❧❦❡s ❡t ❛❧✳ ✷✵✵✽❪✱ ❞❡♥tr❡ ♦✉tr❛s✳

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✸✶

❋✐❣✉r❛ ✶✳✶✿ ❊①❡♠♣❧♦ ❞❡ ♠ás❝❛r❛ ❞❡ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r r❡❢❡r❡♥t❡ à s✉♣❡r❢í❝✐❡ ❞❡ ✉♠ ❡♠❜r✐ã♦ ❞❡ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ❈❛❞❛ ❝♦♠♣♦♥❡♥t❡ ❜r❛♥❝♦ ♥❛ ✐♠❛❣❡♠ r❡♣r❡s❡♥t❛ ✉♠ ♥ú❝❧❡♦ ❝❡❧✉❧❛r✳ ❋✐❣✉r❛ ♦❜t✐❞❛ ❞❡ ❬P✐s❛r❡✈ ❡t ❛❧✳ ✷✵✵✾❪✳

♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s✳ ■ss♦ r❡q✉❡r ♦ ❞❡s❡♥✈♦❧✈✐♠❡♥t♦ ❞❡ ♥♦✈♦s ♠ét♦❞♦s ❝♦♠♣✉t❛❝✐♦♥❛✐s ❡✴♦✉ ❛♣r✐♠♦r❛♠❡♥t♦s ❞♦s ❛t✉❛✐s✳ ❉❡st❛ ❢♦r♠❛✱ ❛♦ ❧♦♥❣♦ ❞♦s ❛♥♦s✱ ❢♦r❛♠ ♣r♦♣♦st♦s ✈❛r✐❛❞♦s ♠ét♦❞♦s ♣❛r❛ ❛ r❡❛❧✐③❛çã♦ ❞❡ss❛ t❛r❡❢❛ ✭♦❜t❡♥çã♦ ❞❛ ♠ás❝❛r❛ ❞❡ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r✮✳ ❆ ♠❛✐♦r✐❛ ❞♦s tr❛❜❛❧❤♦s ❞❡s❡♥✈♦❧✈✐❞♦s tr❛t❛ ❞❡ ✐♠❛❣❡♥s r❡❧❛t✐✈❛s à s✉♣❡r❢í❝✐❡ ❞♦s ❡♠❜r✐õ❡s ❞❛ ❉r♦s♦♣❤✐❧❛ ❬❙✉r❦♦✈❛ ❡t ❛❧✳ ✷✵✵✽✱P♦✉st❡❧♥✐❦♦✈❛ ❡t ❛❧✳ ✷✵✵✹✱❏❛♥ss❡♥s ❡t ❛❧✳ ✷✵✵✺✱❑♦s♠❛♥ ❡t ❛❧✳ ✶✾✾✾❪✱ ❡ ♣❡r♠✐t❡♠ ✐❞❡♥t✐✜❝❛r ✭❞❡ ❢♦r♠❛ ✐♥❞✐✈✐❞✉❛❧✮ ♦s ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s✱ ❝♦♥❢♦r♠❡ ❞❡♠♦♥str❛❞♦ ♥❛ ❋✐❣✳✶✳✷✳ P♦r ♦✉tr♦ ❧❛❞♦✱ ✈❡r✐✜❝❛♠✲s❡ t❛♠❜é♠ s♦❧✉çõ❡s ❝♦♠♣✉t❛❝✐♦♥❛✐s ❞✐r❡❝✐♦♥❛❞❛s à ❛♥á❧✐s❡ ❞❡ ✐♠❛❣❡♥s ♥♦ ♣❧❛♥♦ ❛♥❛tô♠✐❝♦ s❛❣✐t❛❧✱ ❛s q✉❛✐s ❣❡r❛❧♠❡♥t❡ ♣r♦✲ ♣õ❡♠ ❛❜♦r❞❛❣❡♥s ❛❧t❡r♥❛t✐✈❛s ♣❛r❛ ❛ ❝♦♥str✉çã♦ ❞❛ ♠ás❝❛r❛ ❞❡ s❡❣♠❡♥t❛çã♦ ♥✉❝❧❡❛r✳ ❊♠ t❛✐s ❝❛s♦s✱ ♦s ♠ét♦❞♦s ❞❡s❡♥✈♦❧✈✐❞♦s ❣❡r❛❧♠❡♥t❡ s❡ ❜❛s❡✐❛♠ ♥❛ té❝♥✐❝❛ ❞❡ ❥❛♥❡❧❛s ❞❡s❧✐③❛♥✲ t❡s✱ ❡ ♥ã♦ ✐❞❡♥t✐✜❝❛♠ ♣r❡❝✐s❛♠❡♥t❡ ♦s ♥ú❝❧❡♦s ✐♥❞✐✈✐❞✉❛❧♠❡♥t❡ ❬❍♦✉❝❤♠❛♥❞③❛❞❡❤ ❡t ❛❧✳ ✷✵✵✷✱●r❡❣♦r ❡t ❛❧✳ ✷✵✵✼✱❍❡ ❡t ❛❧✳ ✷✵✵✽❪✳

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✸✷ ❈❛♣ít✉❧♦ ✶✳ ■♥tr♦❞✉çã♦

❋✐❣✉r❛ ✶✳✷✿ ❊①❡♠♣❧♦ ❞❡ ✉♠ ❡♠❜r✐ã♦ ❞❡ ❉r♦s♦♣❤✐❧❛ r❡❢❡r❡♥t❡ ❛♦ ♣❧❛♥♦ s❛❣✐t❛❧✳ ❖s ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s s❡ ❧♦❝❛❧✐③❛♠ ♥❛ ❡①tr❡♠✐❞❛❞❡ ❞♦ ❡♠❜r✐ã♦✳

✶✳✶ ❖❜❥❡t✐✈♦s

❖ ♦❜❥❡t✐✈♦ ❞❡st❡ tr❛❜❛❧❤♦ é ❞❡s❡♥✈♦❧✈❡r ♠ét♦❞♦s ❝♦♠♣✉t❛❝✐♦♥❛✐s ❜❛s❡❛❞♦s ❡♠ té❝♥✐❝❛s ❞❡ ♣r♦❝❡ss❛♠❡♥t♦ ❞✐❣✐t❛❧ ❞❡ ✐♠❛❣❡♥s✱ ❞❡ ♠♦❞♦ q✉❡ ♦♣❡r❡ ❝♦♠♦ ✉♠❛ ❢❡rr❛♠❡♥t❛ ❞❡ ❛♣♦✐♦ ❛♦s ❜✐ó❧♦❣♦s ♥❛ ❛♥á❧✐s❡ ❡ ❝❛r❛❝t❡r✐③❛çã♦ ❞❡ ✐♠❛❣❡♥s s❛❣✐t❛✐s r❡❢❡r❡♥t❡s ❛ ❡♠❜r✐õ❡s ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳

✶✳✶✳✶ ❖❜❥❡t✐✈♦s ❊s♣❡❝í✜❝♦s

✶✳ ❉❡s❡♥✈♦❧✈✐♠❡♥t♦ ❞❡ ♠ét♦❞♦s ❝♦♠♣✉t❛❝✐♦♥❛✐s ♣❛r❛ s❡❣♠❡♥t❛çã♦ ❞❡ ✐♠❛❣❡♥s s❛❣✐t❛✐s ❞❡ ♣❛❞rõ❡s ❞❛ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❞❛ ❉r♦s♦♣❤✐❧❛✳ Pr❡t❡♥❞❡✲s❡ q✉❡ t❛✐s ♠ét♦❞♦s s❡❥❛♠ ❝❛♣❛③❡s ❞❡ ✐❞❡♥t✐✜❝❛r ❝♦♠ ♣r❡❝✐sã♦ ❛ ❧♦❝❛❧✐③❛çã♦ ❞♦s ♥ú❝❧❡♦s ❞♦ ❡♠❜r✐ã♦ ❡♠ ❞❡s❡♥✲ ✈♦❧✈✐♠❡♥t♦ ♣❛r❛ ❛ ❝♦♥str✉çã♦ ❞❡ ✉♠❛ ♠ás❝❛r❛✳ ❊ss❛ ♠ás❝❛r❛ ♣❡r♠✐t✐rá ❞❡t❡r♠✐♥❛r ❛s ❝♦♥❝❡♥tr❛çõ❡s ❞❡ ♣r♦t❡í♥❛s ❡ ❘◆❆s ❞❡♥tr♦ ❡ ❢♦r❛ ❞♦s ♥ú❝❧❡♦s✳

✷✳ ❉❡s❡♥✈♦❧✈✐♠❡♥t♦ ❞❡ ♠ét♦❞♦s ❝♦♠♣✉t❛❝✐♦♥❛✐s ♣❛r❛ ❛♥á❧✐s❡ ❞❛s ✐♠❛❣❡♥s s❛❣✐t❛✐s✱ ❝❛✲ ♣❛③❡s ❞❡ ❡①♣❧♦r❛r ❞✐❢❡r❡♥t❡s ♣r♦♣r✐❡❞❛❞❡s ❣❡♦♠étr✐❝❛s ❞♦s ❞❛❞♦s ❜✐♦❧ó❣✐❝♦s✳ ❖s ♠ét♦❞♦s ❞❡ ❛♥á❧✐s❡ ✈✐s❛♠ ✐❞❡♥t✐✜❝❛r ❛ ❞✐str✐❜✉✐çã♦ ❞❡ ♣r♦t❡í♥❛s✱ ❡ ❞❡ss❛ ❢♦r♠❛ ♦s ♣❛❞rõ❡s ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ r❡❣✐str❛❞♦s ♥❡ss❛s ✐♠❛❣❡♥s✳

✶✳✷ Pr✐♥❝✐♣❛✐s ❈♦♥tr✐❜✉✐çõ❡s

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✶✳✸✳ ❖r❣❛♥✐③❛çã♦ ❞❛ ❞✐ss❡rt❛çã♦ ✸✸

❝♦♠ r❡❞✉çã♦ ❞❛ t❛①❛ ❞❡ s✉♣❡r s❡❣♠❡♥t❛çã♦ ♦✉ ❝♦♠♣♦♥❡♥t❡s ❛❣r❡❣❛❞♦s✳

❆ ♣❛rt✐r ❞✐ss♦✱ t♦r♥♦✉✲s❡ ♣♦ssí✈❡❧ ❛♣r❡s❡♥t❛r ✉♠❛ ❛❜♦r❞❛❣❡♠ ♣❛r❛ ❛♥á❧✐s❡ q✉❛♥t✐t❛t✐✈❛ ❞♦s ♣❛❞rõ❡s ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✱ ❝❛♣❛③ ❞❡ ❞❡t❡r♠✐♥❛r ❛s ❝♦♥❝❡♥tr❛çõ❡s ❞❡ ♣r♦t❡í♥❛ ❡ ❘◆❆s ❞❡♥tr♦ ❡ ❢♦r❛ ❞♦s ♥ú❝❧❡♦s✳

❖✉tr❛ ✐♠♣♦rt❛♥t❡ ❝♦♥tr✐❜✉✐çã♦ r❡❢❡r❡✲s❡ ❛♦ ♠ét♦❞♦ ❞❡s❡♥✈♦❧✈✐❞♦ ♣❛r❛ ✈✐s✉❛❧✐③❛çã♦ ❞❛s ✐♠❛❣❡♥s s❛❣✐t❛✐s ❞❡ ❡♠❜r✐õ❡s ❞❡ ❉r♦s♦♣❤✐❧❛✱ ♦ q✉❛❧ ♣❡r♠✐t❡ ❛♥❛❧✐s❛r ♦ ❡♠❜r✐ã♦ ❡♠ ✉♠❛ ❝♦♥✜❣✉r❛çã♦ ❧✐♥❡❛r✳ ❊ss❛ ❝♦♥✜❣✉r❛çã♦ ♣❡r♠✐t❡ ✈✐s✉❛❧✐③❛r ♦s ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s ❞✐s♣♦st♦s ❧❛❞♦ ❛ ❧❛❞♦✱ ❢❛✈♦r❡❝❡♥❞♦ ❛ ❛♥á❧✐s❡ ❞❛ ✈❛r✐❛çã♦ ❡s♣❛❝✐❛❧ ❡ t❡♠♣♦r❛❧ ❞❛ ✐♥t❡♥s✐❞❛❞❡ ♠é❞✐❛ ❞❡ ♣r♦t❡í♥❛ r❡t✐❞❛ ❡♠ ❝❛❞❛ ♥ú❝❧❡♦✳ ❆ ♥♦✈❛ ❛❜♦r❞❛❣❡♠ é ✈❛♥t❛❥♦s❛ ❡ ❝♦♠♣❛❝t❛✱ t❛❧ q✉❡✱ t♦r♥❛ ♣♦ssí✈❡❧ ♠♦str❛r✱ ❞❡ ✉♠❛ só ✈❡③✱ ❞❡③❡♥❛s ❞❡ ❡♠❜r✐õ❡s✳ ❆❧é♠ ❞✐ss♦✱ ♣♦ss✐❜✐❧✐t❛ ❝♦♠♣❛r❛r ♦s ♣❛❞rõ❡s ❞❡ ❡①♣r❡ssã♦ ❞❡ ❞✐❢❡r❡♥t❡s ❡♠❜r✐õ❡s✱ ❢❛❝✐❧✐t❛♥❞♦ ❛ ❛♥á❧✐s❡ ❡ ✐♥t❡r♣r❡t❛çã♦ ❞♦s ❞❛❞♦s✳

❚♦♠❛❞♦s ❡♠ ❝♦♥❥✉♥t♦✱ ♦s ♠ét♦❞♦s ❞❡s❡♥✈♦❧✈✐❞♦s s✉❣❡r❡♠ ✉♠ s✐st❡♠❛ ♣❛r❛ ❛♥á❧✐s❡ ❞❡ ✐♠❛❣❡♥s s❛❣✐t❛✐s r❡❧❛t✐✈❛s ❛ ♣❛❞rõ❡s ❞❛ ❡①♣r❡ssã♦ ❣ê♥✐❝❛ ❞❛ ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✱ ❝❛♣❛③ ❞❡ ♣r♦❝❡ss❛r ❡ ❡①tr❛✐r ♣r♦♣r✐❡❞❛❞❡s ❣❡♦♠étr✐❝❛s ❞♦s ❞❛❞♦s ❛♥❛❧✐s❛❞❛s✳ ❆ ❣❡r❛çã♦ ❞❡ r❡s✉❧t❛❞♦s ❛✉t♦♠át✐❝♦s ❡ ♣r❡❝✐s♦s ♣❡r♠✐t❡ ❡❧✐♠✐♥❛r ❛ s✉❜❥❡t✐✈✐❞❛❞❡ ❞♦ ♣r♦❝❡ss♦ ❞❛ s✐♠♣❧❡s ❛♥á❧✐s❡ ✈✐s✉❛❧✱ ❛❧é♠ ❞❡ ♣❡r♠✐t✐r ❛♥❛❧✐s❛r ✉♠ ❣r❛♥❞❡ ♥ú♠❡r♦ ❞❡ ✐♠❛❣❡♥s ❡♠ ✉♠ ❝✉rt♦ ✐♥t❡r✈❛❧♦ ❞❡ t❡♠♣♦✱ ❝♦♥tr✐❜✉✐♥❞♦ ❛ss✐♠ ❝♦♠ ✈❛r✐❛❞❛s ✐♥t❡r♣r❡t❛çõ❡s ❜✐♦❧ó❣✐❝❛s✳

✶✳✸ ❖r❣❛♥✐③❛çã♦ ❞❛ ❞✐ss❡rt❛çã♦

❊st❛ ❞✐ss❡rt❛çã♦ ❡stá ♦r❣❛♥✐③❛❞❛ ♥♦s s❡❣✉✐♥t❡s ❝❛♣ít✉❧♦s✳ ◆♦ ♣r✐♠❡✐r♦ ❝❛♣ít✉❧♦ ❢♦✐ ❛♣r❡s❡♥t❛❞❛ ❛ ✐♥tr♦❞✉çã♦✱ ♦❜❥❡t✐✈♦s ♣r♦♣♦st♦s ❡ ❛ ♠♦t✐✈❛çã♦ ♣❛r❛ r❡❛❧✐③❛çã♦ ❞♦ ♣r❡s❡♥t❡ tr❛❜❛❧❤♦✳ ❆ r❡✈✐sã♦ ❜✐❜❧✐♦❣rá✜❝❛ ❡stá ❞✐✈✐❞✐❞❛ ❡♠ ❞♦✐s ❝❛♣ít✉❧♦s✳ ◆♦ ❈❛♣ít✉❧♦2❛♣r❡s❡♥t❛✲

s❡ ✉♠❛ ✐♥tr♦❞✉çã♦ ❞♦s ❝♦♥❝❡✐t♦s ❜✐♦❧ó❣✐❝♦s✱ ❡♥❢❛t✐③❛♥❞♦ q✉❡stõ❡s r❡❧❛t✐✈❛s à ❉r♦s♦♣❤✐❧❛ ❡ ❢♦r♠❛s ❞❡ ❡st✉❞♦ ♣❛❞rõ❡s ❞❡ ❡①♣r❡ssã♦ ❣ê♥✐❝❛✳ ❖ ❈❛♣ít✉❧♦ 3 ❞❡s❝r❡✈❡ ♦s ❝♦♥❝❡✐t♦s ❡

té❝♥✐❝❛s ❞❛ ár❡❛ ❞❡ ♣r♦❝❡ss❛♠❡♥t♦ ❞✐❣✐t❛❧ ❞❡ ✐♠❛❣❡♥s✱ ❢✉♥❞❛♠❡♥t❛✐s ♣❛r❛ ❛ ❝♦♠♣r❡❡♥sã♦ ❞♦ ♠ét♦❞♦ ♣r♦♣♦st♦✳ ■♥✐❝✐❛❧♠❡♥t❡ ❛♣r❡s❡♥t❛♠✲s❡ ❢✉♥❞❛♠❡♥t♦s ❜ás✐❝♦s ❞❡ ✐♠❛❣❡♠ ❞✐❣✐t❛❧✱ ❡♠ s❡❣✉✐❞❛ ❛s ❡str❛té❣✐❛s ❞❡ r❡❛❧❝❡✱ s✉❛✈✐③❛çã♦✱ s❡❣♠❡♥t❛çã♦ ❡ ❛♥á❧✐s❡ ❞❡ ✐♠❛❣❡♥s✳ ❖ ❈❛♣ít✉❧♦4 ❛♣r❡s❡♥t❛ tr❛❜❛❧❤♦s r❡❧❛❝✐♦♥❛❞♦s à ❉r♦s♦♣❤✐❧❛ ♠❡❧❛♥♦❣❛st❡r✳ ❖s tr❛❜❛❧❤♦s sã♦

❞✐✈✐❞✐❞♦s ❡♠ três ❝❛t❡❣♦r✐❛s✱ s❡♥❞♦ s✉♣❡r❢í❝✐❡ ❞❡ ❡♠❜r✐õ❡s✱ ♣❧❛♥♦ ❛♥❛tô♠✐❝♦ s❛❣✐t❛❧ ❡ ❜❛s❡s ❞❡ ✐♠❛❣❡♥s✳ ❖ ❈❛♣ít✉❧♦ 5 r❡❧❛t❛ ♦s r❡❝✉rs♦s ✉t✐❧✐③❛❞♦s✱ ❜❡♠ ❝♦♠♦✱ ♦ ♠ét♦❞♦ ♣r♦♣♦st♦

♣❛r❛ ❛ ✐❞❡♥t✐✜❝❛çã♦ ❞♦s ♥ú❝❧❡♦s ❝❡❧✉❧❛r❡s ❛ ♣❛rt✐r ❞❡ ✐♠❛❣❡♥s ❞❡ ❡♠❜r✐õ❡s ❞❛ ❉r♦s♦♣❤✐❧❛✳ ❖ ❈❛♣ít✉❧♦ 6 ❛♣r❡s❡♥t❛ ❛ ♠❡t♦❞♦❧♦❣✐❛ ❞❡s❡♥✈♦❧✈✐❞❛ ♣❛r❛ ❛✈❛❧✐❛çã♦ ❡ ✈✐s✉❛❧✐③❛çã♦ ❞❛ ❡①✲

♣r❡ssã♦ ❣ê♥✐❝❛ ❡♠ ✐♠❛❣❡♥s s❛❣✐t❛✐s ❞❛ ❉r♦s♦♣❤✐❧❛✳ ❖ ❈❛♣ít✉❧♦7❞❡s❝r❡✈❡ ♦s ❡①♣❡r✐♠❡♥t♦s

r❡❛❧✐③❛❞♦s✱ ❛ss✐♠ ❝♦♠♦✱ ❞❡♠♦♥str❛ ❡ ❞✐s❝✉t❡ ♦s r❡s✉❧t❛❞♦s ♦❜t✐❞♦s ♣❡❧♦s ♠ét♦❞♦s ♣r♦♣♦s✲ t♦s✳ ❋✐♥❛❧♠❡♥t❡✱ ♦ ❝❛♣ít✉❧♦8tr❛t❛ ❞❛s ♣r✐♥❝✐♣❛✐s ❝♦♥❝❧✉sõ❡s✱ ❝♦♥tr✐❜✉✐çõ❡s ❡ ♣❡rs♣❡❝t✐✈❛s

(36)

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Segundo Moyers (1991), a perda prematura de qualquer dente decíduo pode facilitar a erupção precoce de seu sucessor permanente, ou retardá-lo de acordo com

Porém, em países onde a legislação para conexão de sistemas fotovoltaicos à rede elétrica permite que o conversor utilizado seja não-isolado, como é o caso do Brasil, em que

Efeitos da congelação lenta e da vitrificação na viabilidade, ultraestrutura e expressão gênica de embriões caninos produzidos