Rantabe, 29.X.1986, 6cT, 4? (L.A.Nilsson et B. Pettersson ; UUDSB)*, Terminalia pumila, ld1, 19, FN (L. A. Nilsson).- Foulpointe, 3l.xii.
1991, Portulacca, 29 ; x.1995, forêt, fauchoir, 19 ; lO.x.1995, cultures, Lobelia agrestis, 1 d ; xi.1995, jardin, 19 (A.Pauly).- Ampasimbé-Onibé, 19.V.1993, banc de sable rivière, ld (A.Pauly).- 46 km N. Brickaville, 8.Ü.1996, Borreria verticillata, 19 (A.Pauly).
DIEGO-SUAREZ : Nosy-Komba, i.1952, 7d, 29 (N.L.H.Krauss ; MNHNP).
MAJUNGA : Station forestière d’Ankarafantsika, 21- 22. xi.1986, 12d, 16 9 (J.Wenzel ; SMUK)*.- 2km E. Mahajanga, 23. xi.1986, 4d, 599 (J.Wenzel; SMUK)*.- Ankarafantsika, Ampijoroa, 22.xi.1986, Erythroxylum platycladum, FN, 19 (L.A.Nilsson & P.Pettersson ; UUDSB)*.
FIANARANTSOA : Isalo, Analalava Malio, lit rivière sablonneuse, 19.iii. 1994, Helinus integrifolius, 79d, 699, Gouania sp., 29d, 249, Indigofera, 38d, Triumfetta rhomboidea, 19 (A.Pauly).- Isalo National Park, 19.iii. 1994, 5 9 (M.Wasbauer ; USUL ; dt Pesenko).
TULEAR : Réserve de Beza Mahafaly, 15.xi.1984, 1 d (R.Brooks ; SMUK)*.- Berenty, 28.ii.1985, 2d, 3 9 (J.Wenzel;
SMUK)*; 5-15.V.1983, 51d, 149 (J.S.Noyes & M.C.DAY. BMNH) ; 3-14.iv.1994, 3d, 89 (M.Wasbauer; USUL; dt Pesenko) .- Toliara 5km N, 22.iii.1994, 4d, 19 (M.Wasbauer ; USUL; dt Pesenko).- Toliara 30km S, 28.xi.1986, 2d, 19 (J.Wenzel; SMUK)*.- Toliara 12km SE, 22-28.iii.1994, 13d ; 12-13.iv.1994, 30d (M.Wasbauer; USUL; dt Pesenko).- 45 km S. Mahabo, 24-26.xi.1986, 19 (J.Wenzel; SMUK)*.- Ankilibe, 22.iv.1984, ld (R.Hensen & A.Aptroot)*.- Betioky, 5.iv.l968, ld (K.Guichard ; BMNH)*.- Betioky 2km N, 24.iii.1994, ld (M.Wasbauer; USUL; dt Pesenko).- Bevilany, 12.iv.1968, 2d (K.Guichard; BMNH)*.- Tongobory, 27.iii.1968, 2d (K.Guichard;
BMNH)*.- St Augustin, 29.iii.1968, ld (K.Guichard; BMNH)*.- Forêt du Zombitsé, 22.iii.1968, 5d (K.Guichard; BMNH).- Behara, iv.1937, 12d, 29 (A.Seyrig; MNHNP)*; 2d, 19 (A.Seyrig; MRACT), xii.1940, 1 d, 39 (ASeyrig ; MNHNP).- Morondava, Bereboka Reserve, 17- 24. v. 1983, 37d, 19 (J.S.Noyes & M.C.Day; BMNH).-
Morondava, lO.xii. 1991, fl. Tamarindus, 2d, Papilionaceae, 12d, Ipomoea pescaprae, ld (A.Pauly).- Ampanihy, 20°-25’S- 44°44’E, 12.xii.1991, fl. 620 = Kochneria madagascariensis, 79 (A.Pau!y).- Belalanda, 23.iii.1994, Euphorbi aceae, ld, Papilionaceae, 5d, 19 (A.Pauly).- Sakaraha 38 Km E, 21 .iii. 1994, 1 d, 29 (M.Wasbauer ; USUL; dt Pesenko).
hyaline ; veins and stigmae pale yellow, except orange-brown vein forming the marginal cells and stigmae. Posterior marginal areas of metasomal terga yellowish translucent.
Vestiture. White, short, not dense, erect, slighltly plumose ; metanotum, sterna and last terga with long liairs ; plumose adpressed hairs absent. »
Variations géographiques. La variété de la falaise orientale est encore plus sombre que la variété du Sud. Les mâles ont le clypeus, le col du pronotum et les pattes (sauf l’intérieur des tibias antérieurs) complètement noirs.
Matériel. MADAGASCAR. TULEAR : Tongobory, 200m, 27.iii.1968, ld (K.Guichard ; BMNH ; Pesenko, 1996).- Betioky 2km N, 29.iii.1994, ld1 (M.Wasbauer; USUL ; dt Pesenko).
TAMATAVE: Didy, 16.iv.1992, forêt, fl. 601 = Acacia farnesiana, 60 d, 9 ? ; Harungana tnadagascariensis, 3 d (A.Pauly).- Morarano-Chrome, 25km W., iv.1992, forêt, fl. 601 = Acacia farnesiana, 4d, 13.iv.1991, fl. 601 = Acacia farnesiana, 13 d, 6 9 ; i.
1992, fl. 643 =Mapouria aegialodes, 5d (A.Pauly).
Ceylalictus (Atronomioides) madagassus Blüthgen
- (Fig. 16, c, d ; PI. 1, O, P)Nomioides madagassa Blüthgen, 1934 : 278, 9. Holotype : 19, MADAGASCAR [TANANARIVE], Antsirabe, 1000-1500m, 27.ix.
1928 (A.Seyrig; col. Hedicke), détruit. Un paratype, idem holotype (MNHUB). Pesenko, 1983 : 108 (N.
madagassus). Pesenko, 1996 (comb.nov.).
Halictus personatus Benoist, 1962 : 108,
<?. Holotype: le?, MADAGASCAR [TANANARIVE], Ambatolampy, 1 .i. 1958 (F.Keiser ; NHMB) ; Pesenko, 1996 (synonymie)
Diagnose. Voir clé.
Distribution. Forêts des plateaux et de la falaise orientale.
Matériel. MADAGASCAR. TANANARIVE : Ambatolampy, l.i. 1958, 2d (F.Keiser; NHMB ; Benoist, 1962).- Ankazobe, Ambohitantely, 17.xi.1983, Weinmannia rutenbergi., ld (L.ÆNilsson et L.Jonsson ; UUDSB ; Pesenko, 1996).- Mt Angavokely, 15.iii.1985, n° 85 :76, 19 (L.A.Nilsson; UUDSB; Pesenko, 1996), sur Emilia citrina, FN (L. A. Nilsson).-
'-■mm'
d
Fig. 18, Ceylalictus (Atronomioides) petiolatus, mâle (a-c) et femelle (d-h).
a, d, tête.- e,f, partie inférieure de la tête d’autres spécimens.- c, cellules submarginales.- g, calcar,- b, h, metasoma.
Fig. 19, Ceylalictus (Atronomioides) sylvestris, mâle (a-h) et femelle (i-k)
a-c, i, tête (vue de face).- d, flagellomères.- e, stemite 7.- f, stemite 8,- g, h, genitalia (g, vue ventrale, h, vue dorsale).-j, calcar.- k, metasoma.
Andriambilany, 31.xii.1957, le? (F.Keiser; NHMB ; Benoist, 1962).
TAMATAVE : Morarano-Chrome 25 km W., forêt, 25.iv.1991, piège bac jaune, 1? (A.Pauly).- Didy, 16.iv.1992, forêt, Mikania ; vi. 1992, piège bac jaune, 1$ (A.Pauly).
Ceylalictus (Atronomioides) aldabranus (Cockerell)
- (Fig. 16, a, b ; PI. 1, M, N)Halictus aldabranus Cockerell, 1912 : 31,
<f. Holotype : ld\ Ile ALDABRA, 27.xii.1908, n°312
= ESPRIT Island (J.C.F.Fiyer ; BMNH) ; Pesenko 1996 (comb.nov.).
Diagnose. Voir clé. Une petite espèce noire à tête ronde.
CeylalictUS aldabranUS 92 spécimens, 32 données
Variations. On distingue deux variétés géographiques à Madagascar.
A. Sur la côte Est (Province de Tainatave), le scutum est mat chagriné à ponctuation fine et dense au centre, les genoux et tarses du mâle noirs.
B. Dans le Sud et l’Ouest (Provinces de Majunga et Tuléar), le scutum est brillant lisse sur les côtés, densément ponctué au centre, le mâle a les genoux et tarses jaunes ; la même variété existe aux Comores et c’est probablement aussi la variété typique d’Aldabra.
Matériel. MADAGASCAR. TAMATAVE : Soanierana Ivongo, N. Rantabe, 28-29.x.1986, le?, 19 (L.A.Nilsson et B. Pettersson
; UUDSB ; Pesenko, 1996), sur Phylohydrax madagascariemis, 1 e?
(L. A. Nilsson).- Soanierana Ivongo, Manambolosy, 27 et 28.X.1986, Lobelia fervens, FN, 29 (L. A. Nilsson).- Foulpointe, 31 .xii. 1991, fl.
630 = Portulacca, 29 ; vii.1993, fl. 630 = Portulacca, 4c?, 2 9 ; l-lO.x.1995, plage, 19 ; x.1995, forêt, 29 ; lO.x.1995, Lobelia agrestis, 59 ; Alternanthera sessilis, 4 c? ; Aptenia cordifolia, 4<? ; 19.xi.1995, fl. basilic = Ocimum basilicum, 3 c? ; xi.1995, fl. 726 = Enhydra sessilis, 3 «? ; fl. 728 = Aptenia cordifolia, 1 <? ; fl. 740 = Launea bellidifolia, le?1; fl. 724 = Aspilia bojeri, 19 ; fl. 730 = Alternanthera sessilis, 3 c? ; xi.1995, plage, piège bac jaune, 1 9 ;
23. xii.1995, Oldenlandia, 4c? (A.Pauly).- Ambila-Lemaitso, lO.x.1971, 19 (L.Blommers ; ITZA).
MAJUNGA : Ankarafantsika, Ampijoroa, 17- 27.xi.1986, 2c?, 3 9 (L.A.Nilsson et B.Pettersson ; UUDSB; Pesenko, 1996), sur Mollugo nudicaulis, FN, 19, Nervi lia crociformis, « seeking », 19, Nervilia petraea, « seeking », 2 c?, 3 9 (L. A. Nilsson).
TULEAR : Morondava, Bereboka Reserve, 17- 24. v. 1983, 6c?, 249 (J.S.Noyes & M.C.Day; BMNH).- Morondava, 50 km E, 9.xii.l992, sur la sueur, le? (A.Pauly).- Toliara 10km NE, 24.iii.1994, le? (M.Wasbauer; USUL; dt Pesenko).-Tuléar, l.v.1971 (L.Blommers ; ITZA).
COMORES. GRANDE COMORE : Le Galawa H., 22.iv-5.v.l991, 4<?, 5 9 (K.M.Guichard ; BMNH).- Irsandra H ;, 23.iv-3.v.l991, 19 (K.M.Guichard ; BMNH).
Ceylalictus (Atronomioides) sylvestris Pesenko & Pauly sp. nov. -
(Fig. 19)Holotype : ld\ MADAGASCAR [TANA- NARIVE], LaMandraka, 2.iv.l991, forêt, fauchoir (A.Pauly col.)
Diagnose. Similaire à C. aldabranus, diffère par les caractères suivants : aire propodéale (metapostnotum) défini ; surface horizontale du propodeum concave (fortement chez le mâle), plus long que le scutellum ; scutum et tergites mats ; pont ventral de la gonobase (des genitalia du mâle) élargi au milieu ; valve pénienne (sagittae) avec de grandes lamelles en forme d’aile.
En outre, diffère des deux formes de C.
aldabranus par le scutum complètement mat à ponctuation très espacée au centre, les maculations jaunes des tergites plus réduites. De la forme B, le mâle diffère par les genoux et tarses noirs.
Description [texte original de Pesenko en anglais].
Mâle.
« Structure. Lenght 4.2-4.4 mm. Head flattened, rounded in frontal view ; as high as wide
or slightly higher. Face strongly impressed on the level of antennal sockets. Ocellar élévation distinct Médial clypeal lobe weakly convex ; its height/width ratio 0.95. Clypeus projecting 0.55- 0.60 of its height below lower inargin of eyes.
Supraclypeal area convex, before its upper border with veiy small rounded tubercle. Malar space linear.
Inner orbits with relatively deep triangular notch ; its depth about 0.4 of maximal (extrapolated) ocular width in frontal view; paraocular area in the notch distinctly projected as shiny tubercle.
Longitudinal carina between antennal sockets absent.
Frontal line indistinct. Mandibles with subapical dens.
Antennae relatively long, reaching the end of mesosoma. First flagomere 2.5 times shorter than second one ; second flagomere 1.2 times shorter than subséquent ones, their lenght/diameter ratio 1.2.
Scutellum convex. Metapostnotum crescent- shaped, distinctly impressed, bordered laterally and posteriorly by distinct change of sculpture and rounded élévation. Horizontal surface of propodeum slightly (1.1 times) longer than scutellum, forming with posterior vertical surface a rounded angle of about 135’. Marginal cell of forewing relatively narrow, very narrowly rounded at distal end ; second marginal cell trapezoidally triangle (almost triangle).
Hind wings with six distal hamuli on anterior inargin.
Metasoma moderatly convex, inversely lanceolate, metasomal tergum 6 enlarged, broadly rounded at posterior end. Postgradular areas of terga 2-4 strongly convex. Posterior marginal areas of terga flattened, distinctly separated from postgradular areas ; ones of terga 1 and 2 narrow, ones of terga 3 and 4 relatively broad. Posterior margins of metasomal stema 2-5 straight. Posterior margin of metasomal sternum 7 strongly protuberant medially. Apical lobe of sternum 8 triangularly trapezoidal (fig. f). Génital foramen rounded ; médian gonobase suture présent partly ; posterior margin of gonobase bow- shaped on dorsal surface ; ventral gonobasal bridge moderatly broadened in the middle, situated behind ventral gonocoxal bridge ; gonostyli trapezoidally broadened apically, relatively simple, hairless ; pénis valves broadened medially, with large rounded wing-shaped latéral laminae.
Sculpture. Clypeus dully shiny, excepting shiny upper fifth, obscurely granulate with several raunded pits on dark parts. Supraclypeal area silk- dull, densely superfïcially finely granulate. Frons and vertex dull, densely granuloso-punctate. Gênai areas shiny, obscurely striate, with not dense, irregular and indistinct punctation on upper part behind eyes.
Scutum silk-dull, uniformly densely granulate (each granula equal to eye facet). Scutellum shiny, with sparse fine variable punctation, interspaces polislied.
Metapostnotum
dull, with dense fine longitudinal ruguloses throughout or only laterally and granulate medially.
Adjaccent parts of horizontal surface propodeum and its latéral surfaces dull, densely and very finely granulate ; its posterior vertical surface dull, granulate, with several strong oblique rugoses. Mes- and metepisterna silk dull, superfïcially, finely and densely granulate. Metasomal tergum 1 and postgradular area of terga 2 and 3, densely granulate, these area of suceeding terga becoming shiny ; posterior marginal area of terga shiny, obscurely transverse striate.
Coloration. Black without metallic tints.
Pale integumental marking poor. The following parts whitish : médian spot on labrum ; variable anchor-shaped figure on clypeus; posterior lobes and spots on collar of pronotumlar crests (sometimes partly) ; médian part of metanotum ; pro- and mesobasitibial plates ; very narrow and interrupted medially transverse bands on posterior part of pregradular area of metasomal terga 2 and 3 or only tergum 2. Scapi and pedicels black ; flagella dark brown on lower surface and black on upper surface.
Wing membranes and tegulae distinctly infiiscated ; tegulae, veins and stigmae dark brown. Legs dark brown. Posterior marginal area of metasomal terga brownish translucent.
Vestiture. White, short, not dense, erect, slightly plumose ; relatively long arround antennal sockets, on vertex, gênai areas, and metanotum.
Plumose adpressed hairs absent. » Femelle.
« Structure. Lenght 4.8-5.0 mm. Head flattened, rounded in frontal view ; its height/width ratio 0.94-0.98. Face distinctly and broadly impressed laterally and above antennal sockets. Ocellar élévation distinct, but weak. Médial clypeal lobe flattened, its height/width ratio 0.6. Clypeus projecting 0.75 of its height below lower margin of eyes. Supraclypeal area convex. Malar space linear. Inner orbits with weak rounded notch ; its depth about 0.2 of maximal (extrapolated) ocular width in frontal view; paraocular area in the notch fiat. Longitudinal carina between antennal sockets strong. Frontal line developped.
Scutellum convex. Metapostnotum triangularly crescent-shaped, slightly impressed, bordered by change in sculpture and weak élévation, medially reaching to posterior vertical surface of propodeum.
Horizontal surface of propodeum as long as scutellum, formings with posterior vertical surface a rounded angle of about 135’. Wing venation similar to that in male, excepting variable number of distal hamuli of hind wings (6-7). Inner metatibial spurs with three teeth. Metasoma flattened, broadly inversely lanceolate; postgradular areas of metasomal terga 2-3 broadly convex, posterior marginal areas of
terga relatively broad, distinctly separated from postgradular areas by dépréssion.
Sculpture. Clypeus polished on pale parts ; its dark parts and supraclypeal area densely and very finely granulate with several superficial pits. Frons and vertex dull, granuloso-shagreened. Genal areas silk-shiny, punctate on upper part behind eyes and longitudinally distinet striate near probscidial cavity.
Sculpture of meso- and metasoma similar to that of male, excepting granulate (without rugose) posterior vertical surface of propodeum, duller metepisterna, dull and granulate of posterior marginal area of metasomal terga 1-3.
Coloration. Similar to that of male, except as follows : labrum without yellow spot ; pronotal corolla with yellow band interrupted medially ; posterior part of postgradular area of metasomal terga 2-4 or 2-5 with narrow yellow band interrupted medially and broadened laterally.
Vestiture. Similar to male. Metasoma with developed scopa formed by whitish long, slightly plumose hairs on postgradular area of sterna 2-5. »
CeyllllictUS sylvestris 8 spécimens, 5 données
Distribution. Forêts de la falaise orientale, où elle semble remplacer C. aldabrcmus.
Paratypes : TANANARIVE : idem holotype, 3d, 1$
(A.Pauly réc. ; 2 d col. Pauly, 1 d col. ZISP)
T AM AT AVE : Périnet, 1.xi. 1995, forêt, fauchoir, 19 (A.Pauly).- Morarano-Chrome 25 km W, xi. 1991, forêt, bac jaune, 29 (A.Pauly).
Ceylalictus (Atronomioides) rostratus Pesenko
- (Fig. 16,
e, f ; PI. 1,
Q, R )Ceylalictus (Atronomioides) rostratus Pesenko, 1996: 506, c??. Holotype: ld\
MADAGASCAR [TAMATAVE], Soanierana Ivongo, N. Rantabe, 29.x. 1986 (L.A.Nilsson et B.Pettersson ; UUDSB).
Diagnose. Une espèce noire à tête remarquablement allongée.
Distribution. Littoral Est de Madagascar.
Fleurs butinées. Nombreuses fleurs sur la plage, en particulier sur Scaevola spp.
(Goodeniaceae).
Matériel. MADAGASCAR. TAMATAVE : Soanierana Ivongo, N. Rantabe, 28-29.x. 1986, Enhydra sessilis, FN, 3 9, Phylohydrax tnadagascariensis, FN, 4 c?, 19 (L. A Nilsson) .- Vohitsara, 31.x. 1991, plage, 8<?, 79 (APauly).- Foulpointe, 2.xi.l991, plage, bac jaune, 3<?, 19 ; 31.xii.1991, plage, fl. 631 = Scaevola taccada, 1 <?, 19 ; fl. 630 = Portulacca, 5 s , 69 ; fl. 632 = Launea bellidifolia, le? (APauly).- Foulpointe, embouchure de l’Onibé, 9.V.1993, 14c?, 129 sur Ipomoea pescaprae, Launea bellidifolia, Phyllohydrax madagascariensis, Enhydra sessilis; 10.xi.1995, 89c?, 259, sur Scaevola taccada et Scaevola plumieri; xi. 1995, fl.739 = Phylohydrax madagascariensis, 5 9 ; fl. 726 = Enhydra sessilis, le?,39 (APauly).
FIANARANTSOA : Mananjary 2km S, 5m, 13.iv. 1998, beach végétation, Malaise trap, 1 <?, 19 (M.E. Irwin & E.I.
Schlinger ; CAS).
Ceylalictus (Atronomioides) tumidus Pesenko -
(Fig. 16, g, h;
PI. 1, S)
Ceylalictus (Atronomioides) tumidus Pesenko, 1996: 509, c ? ? . Holotype : ld\
MADAGASCAR [TULEAR], Tuléar, 24.xi.1984, fleurs de Parkinsonia aculeata (R.W.Brooks ; UUDSB).
Diagnose. Voir clé. Proche de rostratus mais tête moins allongée.
Distribution. Sud et Ouest de Madagascar, où elle remplace rostratus.
Fleurs butinées. A rechercher dans Ipomoea pescaprae (Convolvulaceae) sur la plage. Butine aussi Parkinsonia aculeata.
Matériel. MADAGASCAR. TULEAR : idem
holotype : le?, 29 ; idem, Ipomoea pescaprae, 89 (R.W.Brooks ; SMUK & ZISP ; Pesenko, 1996).- Ankilibe, 22.iv.1984, le? (R.Hensen
& AAptroot ; RMNH ; Pesenko, 1996).- District de Tuléar, Haut Anakao, 7.iv,1953, 4c? (AR. ; MNHNP).- Morondava, 10.xii.1991, plage, Ipomoea pescaprae, 6 s (APauly).- Tuléar, 23.iii.1994, le?
(APauly).- Ifaty 11km S., 22.ix.1993, sweep of flowering desert scrub, 2c? (W.E.Steiner & R. Andriamasimanana ; USNM). - Ifaty, 30m, 18.iv.1998, hand netted at beach dunes, 49, le? (M.E. Irwin & E.I.
Schlinger; CAS).