Relações morfológicas e morfométricas entre Litopenaeus vannamei (Boone, 1931) e Litopenaeus schmitti (Burkenroad, 1936) (Crustacea, Decapoda, Penaeidae) e suas implicações sobre bioinvasão, no litoral oriental do nordeste do Brasil

Texto

(1)

(2) . E E. 23456789 7

(3) 4 874 4374 8 37 4 8 87 484!"#$%"!!"& '6. 4()*+),4 -!"#$$./&'608148 7

(4) ()*+2,'380 747(4475

(5) 7(6474

(6) 74,4 07 5374 9849 77 8 ( 3 873 8473

(7) 8

(8) 4 4

(9) 687 3 4 4878

(10) 4 4 87

(11) 7584 47

(12) 77 68 877

(13) 4 6 9:87

(14) 078 4; 4<74=7 308 (

(15) 348

(16) 46 >7

(17) 7?748

(18) 7

(19) 4@4

(20) 4873

(21) :87

(22) 4

(23) A 84( 5784

(24) 84B0 5787 948

(25) C03

(26) 4 4 844; 4<74=7 308 D E E F847

(27) 8G68 D48D@H37 2083

(28) 4 87 @8484 3 9 847

(29) 8G68 D48D;8 77I048 J7 3 9 847

(30) 8G48D3783 =

(31) 078

(32) K740784

(33) 423478 A2L29A MN)O.

(34) DEFGHIJFKLKHMNHKHILOMKPMQFPMRILEKHMKPMSPITHMUMDNQS VFJTHWLMKHMXFYOFPTHZLJMUMV[VX[ \LTLOP]L^_PMKHM`aYOFZL^_PMELMNPETHbMDNQSMUMXFYOFPTHZLMVHTPIFLOM`IPcbMdHPePOKPMSHOJPEMUMU\HETIPMKHMXFPZFfEZFLJMUM\X 0123456894 32

(35) 153 4 49345121935412123542414345 334 1146 !" #4143455$ 2%4213 6 !"&# '253436(431 36)434 34#619123912439 1 *12 454 1 2359+894 32

(36) 153 4012345,*3396-. / !09 (554231104523 1#,2425 3 434 4239 11423 4 1*124'421 4 1535)21233 4)5,423 3114 65473481911 9243 120)21(23:918291 40123532424 '11243 120)21(2'329158 32 11$3(324 4 89432 '11243 120)21(2621033;421<3 8=923,(5542311-85435123, (5542311"01214234123,(5542311> 6932 3 412193,(5542311?3242463:91 8291 401235??894326'329158 32 11$3(324 4 ???36621033;421<?@2425 3 434 4239 11 423 4 1*124@AB91 *+23+ 68,' '(2&"!C -.

(37) 23456789 7

(38) 4 874 4374 8 37 4 8 87 484!"#$%"!!"& '6. 4()*+),4 -!"#$$./&'608148 7

(39) ()*+2,'380 747(4475

(40) 7(6474

(41) 74,4 07 5374 9849 77 8 ( 3 873 8473

(42) 8

(43) 4 4

(44) 687 39 4 4878

(45) 4 4 87

(46) 7584 47

(47) 77 68 877

(48) 4 6 :;87

(49) 078 4< 4=74>7 308 (

(50) 348

(51) 46 ?7

(52) 7@748

(53) 7

(54) 4A4

(55) 4873

(56) ;87

(57) 4

(58) B 84( 5784

(59) 84C0 5787 948

(60) D03

(61) 4 4 844< 4=74>7 308 9 . 62B32>E2 FB24G2 . HHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHH 68 9489AI37 2083

(62) 4 87 A8484 @748

(63) 7

(64) 4A4

(65) 4873

(66) ;87

(67) 4

(68) B 84J@AB . HHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHH 68 K948K9477214<L M71 4

(69) 4 @748

(70) 7

(71) 4A4

(72) 48730873

(73) 4648790 J@A6> . HHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHHH 68 K948K9077<40872

(74) 87

(75) 4 @748

(76) 7

(77) 4A4

(78) 4873

(79) N8O03 48 . B773 PQ)R.

(80) . 3. 12345628929

(81) 6. 6828 482

(82) 89462 4 52 2 4 2868 1 42 2 !"#$%!&'%!&(!(#)*&!''!+$,"-#.%#(##'/#01'2 3.

(83) 1234467829 4

(84) 9 28627 4. !"6 #192692$%&'($'($%)*) +,-.%/+%,&%&+000)*+)$+.0%&/+&1/ )+1///+%/%/'(%+20)+0$.'+-3+% )+1)%)4)0///%%//$+4/-5 /,1)+160)&1%+600,//+1/%7%//)-/ /+0/89/:;9

(85) 6 1<=6 > 9! 4? 3 9 @A#!278 9$%& )'$)/&1/$)+B/$%& %)0%'( )+ +1 +'( %//& %//&$%)%&+$%&+,5)*/-.%/+ /+% /%/'(%/+1/C5/)3/ > 742!D E 86 7 ! 2A2427" !"627 >627FG68 2H287 !I

(86) 68 J16764;96 K248 L8M6NOPQRQPSNTQUT9RV%&+%260%+0)&/ &&WXM0.+/+0/) >. 9 27 YZ 2#[29G268 27 2K244 ! 2EF"2!:3[296 9L8%/V%&-&/ /0/))0%\+/ ]7463 ^9 46!269 126 #_62722 4@28394 4E 39 64@27 "`"264 LabOV%&))//(&)'0&,%-'(+&-&7.) c76"2946 22 29 ! @6 d9 7 2 E 92$KI4ed9 3 YZ 2:642`68 2 ?" !3YZ $A2[ 9 27 2^ f768 2g. !

(87) 6 LhbiV . 4 44234G3786 7`96 4 %/00$+/)'j/-&1/%-&,/k+ /%)&$ .'0)Bd642!2$7/9X0'($%&'(/%/%&//// 40/5/)5///-//%)+$.//60,/%.

(88) 1234567897

(89)

(90) 39

(91) 6

(92) 3 !"#$!"%7

(93) 3%3&7'6

(94) 3()"$% *$+%,7+6%3&%-7%+6

(95) &8

(96) %..%79/,768967&%.071

(97) 6787792%63878%% +%,

(98) 3.

(99) .%&

(100) 382%6&8

(101) 3 345446)""79%.3&6767. 83+,737

(102) ,

(103) '

(104) 87.071

(105) 6787 7788.7%

(106) &699:67.+767

(107) .%98%3%2

(108) ,2.%&

(109) +6

(110) 33

(111) 7,

(112) 316

(113) 3 ;63 <=!4>?4<"$$!<=!4@?4A%B=4A?4<"+%,

(114) 3

(115) .%&6

(116) 3% 39'%3&C%38967&%7%,79

(117) 67

(118) 8

(119) +6

(120) 1%&

(121) 8%+%37937

(122) 316

(123) 3 ;63 3!4>$!4 D=E%F$G43?4?4""%7

(124) ;

(125) 9&

(126) 6@"H4D?43?4>+%,

(127) 3

(128) 3%,0

(129) 339'%3&C%3 6:&73%

(130) 66% C%37I778%797,7

(131)

(132) 16

(133) 3$$J!4K?4K!E"+%,73+76%6738%&6797,0

(134) +

(135) 6%3&763%.+6%83+

(136) :2%, 7719876%7

(137) &6996 L9&

(138)

(139) 96'78

(140) +%,73

(141) 2%6373+%,

(142) &%6M.9

(143) +6

(144) 33

(145) 7,%+%,7 7.-78%

(146) A=645"4#=54"4#$H4"4$H45"43!43N=O5PPQR+

(147) 6 &

(148) 87767%3&69&967%.7&%67383+

(149) 9,-78

(150) 3

(151) 3&%'67&%38

(152) 5PPQ3

(153) 9

(154) 6%&7

(155) 8

(156) 16

(157) QS,2

(158) Q6%6%T !"$"4>?4U?45"4">$4#5=54(?4?43$"!V"$!4?4>?4 !WXY4?4$E"D*4K?4V"!%3!4>?4>G$E L9&

(159)

(160) 96'78

(161) +%,7 7.-78%6%3+%&

(162)

(163) 7 7

(164) 2287&6

(165) 78%8Z73+%,771987%.&

(166) 8

(167) 3

(168) 3&6797,0

(169) 3 +%,

(170) 3%1[

(171) 3

(172) 3976

(173) 3+%,7382%6&873

(174) ,%&73

(175) &9673+%,73+737'%3.7672,0

(176) 37379%

(177) .+76&,07.

(178) 3+%,73

(179) 2%6373 % +

(180) 6 &

(181) 8

(182) 3

(183) 3 .

(184) .%&

(185) 3 79% &2%.

(186) 3 19&

(187) 3 \

(188) .+,%.%&76.%&%7UG43?4U?4V?4"]!"4A?4<]$Y4]$">$!4 K=66!%E"=43E= L9&

(189)

(190) 96'78

(191) +%,

(192) 379/:,

(193) 373

(194) ,%&739

(195) .%&673%&7.9Z. +%,77.-78%%6%3+%&

(196) 78779%%8.%67.

(197) 12345678389 5

(198) 79 5342 72

(199) 7

(200) 82457971 87 !"#$ %$ &'()* %*'+,$- %.$(/" 0"*-+ )*1."2"' 01+3$45 6".* /"7+- 5 38-9:8 2 2

(201) 8 742 ;5<7 29=8=>987;7

(202) 5

(203) 875;5<7329=5

(204) 3734248774879 123768 2382< 2345

(205) 747?@226237837894726;7

(206) 8<A268 2362659 23 89

(207) B=58395337689A7C2

(208) 9747774>687-8$** )$#$D'+ E'" F"$ 3D"(2 !"""+++ /$G$#+ 3$*1'$/$+ %$H" 8$$*$&$"%$H",I$"1$52.++,J,"#9 12365353 867423768 2345<29 747 7%$K%L&'(+%$(+K%$M'*L 53G$--N1989A7O9P585;233229 7

(209) 26=2>3;

(210) 72545

(211) 5=85

(212) 9 123653;783/I+*Q&R+5%$GH($56538

(213) 6@23&'(53""9123 456783656

(214) 236783;

(215) S862347689A7

(216) 7945T76B<873EM'"U"*$&-3 &'HV E"-%$+3-#%$+5EQ8+(9P5627;2825;

(217) 2 5?@245=2>3 974797689A7=84735

(218) 87;233B=5<9 1 689A767

(219) 7=8<A23753;237 768 7 26;79A58

(220) 75 76W6 2<5 7;

(221) 2T8338297< X"GJ/J#%$+9Y

(222) 8 742;2

(223) 657;287

(224) 3;2

(225) 7

(226) 57C47

(227) 56 24232362659 23 453457

(228) 747?@27 W2T864533748335

(229) 7?@2;

(230) 2=7=5<659 57 W2T8642392332348739Z5 76268 29 123;53742

(231) 53)#1 &' 6"R )" ["V'58\ ;2

(232) 24223;2

(233) 5 26;5 >987545487?@24

(234) 79 5732<5 7391

(235) 745?268 27768]745572<A8659 245. 24239Y

(236) 8 742;2

(237) 7 59 7

(238) 92332359C^235;2

(239) 9232T5

(240) 55

(241) ;58S5345<882323535<53 3756T7]5

(242) 9.

(243) 1234567829

(244) 484532944886246425494648294589747 624732247997847 1! 945325"4342492#462$%62841697&#1'()*+, 329 54394259464-49.2-475577/04582#1 15 19224552975 322975 4!556786986:!;< 4 =>? 5!569867!;@5; 347 8532-87848497+773797784245784557855497/.2. 6. 6. :ABCD6D4BE7FDG6.

(245) 123456. 8

(246)

(247)

(248)

(249) . !

(250) " #

(251) . $ !%

(252) !

(253) . $ &

(254) '

(255)

(256) &

(257) " # &! '

(258)

(259) (

(260) $ .

(261) !

(262) ! )!% $

(263)

(264)

(265) &! !

(266)

(267) ! . $ *

(268) " +

(269)

(270) )! )

(271)

(272) . ,%

(273) -./0123425678433492. $

(274) & ! -:76;<9.//.

(275) $ '! & )! ,

(276)

(277) , ' ) ) !

(278) % ," # ! )!%

(279) , ' .

(280) ) (

(281)

(282) '

(283) $

(284)

(285) .

(286) ()

(287)

(288)

(289) =

(290) )> " #!$

(291) &

(292)

(293) !

(294) ) !

(295) % , . .

(296)

(297) .

(298) !

(299)

(300)

(301)

(302) $ .

(303)

(304) $

(305) " +,

(306)

(307) &$ ) & &

(308) =

(309)

(310) $

(311) -:78433492.7

(312) ? . " @' ! ) ) !

(313) % , )

(314) )!% .

(315) $

(316) $

(317) )

(318).

(319) ) !

(320) % , . $

(321)

(322) $ &

(323) () ) ( !

(324) $ -:78433492. !

(325) ! !

(326) &

(327)

(328) !" . ABCBDEB3FGHBD2I )

(329) $

(330) $ !

(331) ! . )!% .

(332) 12345674. 9

(333)

(334)

(335)

(336)

(337)

(338)

(339)

(340)

(341)

(342)

(343)

(344)

(345)

(346)

(347)

(348) !"

(349) 9

(350) !

(351)

(352)

(353)

(354)

(355) !

(356)

(357)

(358)

(359)

(360)

(361)

(362)

(363)

(364) !

(365) #!

(366)

(367)

(368)

(369)

(370)

(371)

(372) $ "

(373) %

(374)

(375).

(376)

(377)

(378)

(379) $

(380) !

(381) &

(382)

(383)

(384) ! $

(385)

(386) !

(387) !

(388)

(389)

(390)

(391)

(392) $

(393)

(394)

(395)

(396) '

(397) "

(398) 9

(399)

(400)

(401)

(402)

(403)

(404)

(405)

(406) (

(407) )*+,-./0.12340//05.*

(408)

(409) 67

(410)

(411) &

(412) )8329:5*++*&

(413) $

(414) (

(415)

(416)

(417)

(418) (!

(419)

(420)

(421)

(422)

(423)

(424) "

(425) 9

(426)

(427) !

(428)

(429)

(430)

(431)

(432) (!

(433)

(434)

(435)

(436) $

(437)

(438)

(439)

(440)

(441)

(442)

(443) $

(444)

(445)

(446) "

(447) ;

(448) &

(449)

(450) !

(451)

(452)

(453)

(454)

(455)

(456)

(457)

(458)

(459) (

(460)

(461)

(462)

(463)

(464) !!

(465)

(466)

(467)

(468)

(469) $

(470)

(471)

(472) $

(473)

(474) ! "

(475) 9

(476)

(477)

(478)

(479) <

(480)

(481)

(482)

(483) 7

(484)

(485)

(486)

(487) !

(488) $

(489)

(490)

(491)

(492) $!

(493)

(494) $

(495)

(496) )83 40//05.*

(497) !

(498)

(499) !

(500)

(501) "

(502) ;

(503)

(504) !

(505)

(506)

(507)

(508)

(509)

(510) (&

(511)

(512)

(513)

(514) $

(515)

(516)

(517)

(518) &

(519)

(520)

(521)

(522)

(523)

(524) $

(525)

(526) (

(527)

(528)

(529)

(530)

(531)

(532) &

(533) $!

(534)

(535)

(536)

(537)

(538)

(539)

(540)

(541) $

(542)

(543) !

(544)

(545) )8340//05.*

(546)

(547) !!

(548)

(549)

(550)

(551)

(552)

(553) !

(554)

(555)

(556) =>?@A5B3C

(557)

(558) &

(559)

(560) &

(561) #!!! &

(562)

(563)

(564) .

(565) 123457892

(566) 53. 9 !"#$%&'()&*+,-./01234(5()6*(57%(89:;<*,+=;5:+>+:(&+>,( %8(?(8@>+&+,%>:*3AABA*8@&;8():*,*A(C+>*:D&+?3$+&&+,((5<+>+B.E883 #F*,;C;<+,*,(A6+)-.//G2333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333HI 9 JK(&8;)*>*L;+5(<+&+<:(&M5:;<+5,+L();:9>;+(?:(&)+-@(:+58+#N2(:O>;<*#$2 %:;>;P+,+5)+:+?*)*8;+,(<+8+&Q(5R()+(*;,(+3F*,;C;<+,*,(RO&(PST+&C+):(#./1/23 3333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333H1 9 UR(:+58+,(5@@3(8,;C(&():(58*8():*5*):*L()O:;<*5-,(V*=(8+ +,%>:*3NS4-!",($+;+T*&8*5+SWXYZS[-\"- C+P(),+,(<%>:;=*3];5^*,*&5+>3$+&&+5,((5<+>+B.883$4ZB$*&,+,;5:+>(?:(&)+ ,*>D_%>*>+:(&+>YR``BR&*V(aQ(5,+&(L;^*(?:(&)+,*>D_%>*>+:(&+>YN`FBN_+,* >D_%>*8(,;+)*3333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333330E 9 bKO>;<*,(5@@3(8,;C(&():(58*8():*5*):*L()O:;<*5-,(V*=(8+ +,%>:*3NSA-!",($+;+T*&8*5+SWXY4ST-\"- C+P(),+,(<%>:;=*3$+&&+5,((5<+>+B.883`NFB`+8(>+5(8;S<;&<%>+&,*(5:(&);:* cdddYRFBR>+<+8(,;+,+)*(5:(&);:*cdddYAN`BA&;5:+e):(&*S>+:(&+>,*(5:(&);:* cd]YR&FBR&*:%_(&e)<;+8(,;+)+,*(5:(&);:*cdddYRRABR*&a^*@&*?;8+>,+<*?+,* 7%+&:*@+&,(@(&(;D@*,*5YZfBZ>(=+aQ(5*_>M7%+5,*(5:(&);:*cd]3333333333333333333333333330H 9 gR(:+58+,(5@@3,+<>+55(,(:+8+)6*H3NS$-!"- h#AAi.j-k882-$+;+T*&8*5+SWXYAS4-\"-h#AAi.H-GG882- C+P(),+,(<%>:;=*3];5^*,*&5+>#N(A2(=():&+>#$(42,+8(:+,((57%(&,+3$+&&+,( (5<+>+iNS$B.88YAS4BE-I88333333333333333333333333333333333333333333333333333333333333333333333333333333333j. 9 lR(:+58+,(5@@3,+<>+55(,(:+8+)6*03NS$-!"- h#AAiHk-j882-$+;+T*&8*5+SWXYAS4-\"-h#AAi.0-GE882- C+P(),+,(<%>:;=*3];5^*,*&5+>#N(A2(=():&+>#$(423NS$8(:+,(,;&(;:+YAS4- 8(:+,((57%(&,+3$+&&+,((5<+>+iNS$B.88YA(4BE-I8833333333333333333333333333333333333j0 9 mR(:+58+,(5@@3,+<>+55(,(:+8+)6*j3NS$-!"-h #AAi..-/882-$+;+T*&8*5+SWXYAS4-\"-h#AAi.1-H0882- C+P(),+,(<%>:;=*3];5^*,*&5+>#N2(=():&+>#$2,+8(:+,(,;&(;:+Y=;5^*,*&5+>#A2( =():&+>#42,+8(:+,((57%(&,+3$+&&+,((5<+>+iNS$B.88YAS4BE-I8833333333333333jI 9 nR(:+58+,(5@@3,+<>+55(,(:+8+)6*I3NS4-!"- h#AAiH.882-$+;+T*&8*5+SWXYZST-\"-h#AAi./-EG882- C+P(),+,(<%>:;=*3];5^*,*&5+>#N(Z2->+:(&+>#$2-=():&+>,+@*&a^*,;5:+>+8@>;+,+ #A2(=():&+>#4(T2,+8(:+,((57%(&,+3$+&&+,((5<+>+iN-$(4B.88YABE-HI 88YZSTBE-I88333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333333jk 9 oR(:+58+,(5@@3,+<>+55(,(:+8+)6*13NSA-!"-h #AAiHj-E/882-$+;+T*&8*5+SWXY$S4-!"-h#AAiHI-EG882-$+;+ T*&8*5+SWXYZST-\"-h#AAiHj-j1882-C+P(),+,(<%>:;=*3];5^* ,*&5+>#N2-,*&5+>+_(&:*#$2-=():&+>#A2(=():&+>+_(&:*#42Y=;5^*,*&5+>#Z2(=():&+> #T2,+8(:+,(,;&(;:+3$+&&+5,((5<+>+B.883333333333333333333333333333333333333333333333333333333333333333j/ 9 pR(:+58+,(5@@3,+<>+55(,(:+8+)6*k3NS$-!"- h#AAiH1-1882-$+;+T*&8*5+SWXYAS4-\"-h#AAiHI-EH882- C+P(),+,(<%>:;=*YZST-\"-h#AAiHk-./882-C+P(),+,(<%>:;=*3 ];5^*,*&5+>#N-A(Z2-(=():&+>#$-4(T23];5^*,+8(:+,(,;&(;:+#A(423$+&&+,( (5<+>+iNS$BH88YASTB.883333333333333333333333333333333333333333333333333333333333333333333333333333333333333I..

(567) 123456889

(568)

(569)

(570)

(571) !" # $%&' ()*+'. , -../ 0#'0 1' &2 3"4"%56 .%7' (8+' , -../ 09':; 1'. <=

(572)

(573) > 2?"@ A2B" "4 -$

(574) .1'

(575) ?

(576) 4 -&

(577) 71 A2B"

(578)

(579) 24

(580) 2 -71. &44

(581)

(582) / $%& C 0 @ .%7 C D EF. 1234568G9

(583)

(584)

(585)

(586) !" 9. $%&' ()*+'. , -../ FH'9 1' &2 3"4"%56@ .%7' (8+' , -../ F0'DD 1'. <=

(587)

(588) > 2?" A2B" "4 -$

(589) .1'

(590) ?

(591) 4 -&

(592) 71 &44

(593)

(594) / $%& C 0 @. .%7 C D E:. 1234568I9

(595)

(596)

(597)

(598) !" DH $%&' ( 8+' , -../ :H'0E 1' <=

(599)

(600) > 2?" A2B" "4 -$1

(601) ?

(602) 4 -&1 &44.

(603)

(604) C D EE. 1234568J9 KL2"

(605)

(606)

(607) !" D (8+' M. -../ DH 1' <=

(608)

(609) > 2?" A2B" ?

(610) 4 &44

(611)

(612) C D E;. 1234568N9 KL2"

(613)

(614)

(615) !" 0. $' ()*+' M. -../ D0'O 1' &2 3"4"%56@ &' (8+' M -../ D0';F 1'. <=

(616)

(617) > 2?" $%&' ?2B" ?

(618) 4 &44

(619)

(620) C D E#. 1234568P9 KL2"

(621)

(622)

(623) !" F. $' ()*+' M. -../ DE'0 1' &2 3"4"%56@ &' (8+' M -../ DF'O0 1'. <=

(624)

(625) > 2?" $%&' ?2B" ?

(626) 4 &44

(627)

(628) C D ;H. 1234568Q9 KL2"

(629)

(630)

(631) !" :. $' ()*+' M. -../ DO'E 1' &2 3"4"%56@ &' (8+' M -../ D#'HO 1'. <=

(632)

(633) > 2?" $%&' ?2B" ?

(634) 4 &44

(635)

(636) C D ;0. 1234568R9 KL2"

(637)

(638)

(639) !" E. $' ()*+' M. -../ 0H 1' &2 3"4"%56@ &' (8+' M -../ D9'F9 1' <=

(640) .

(641) > 2?" $%&' ?2B" ?

(642) 4 &44

(643)

(644) C D ;:. 1234568S9 KL2"

(645)

(646)

(647) !" ;. $' ()*+' M. -../ 0F'F 1' &2 3"4"%56@ &' (8+' M -../ 00'O9 1'. <=

(648)

(649) > 2?" $%&' ?2B" ?

(650) 4 &44

(651)

(652) C D ;;. 123456GT9 KL2"

(653)

(654)

(655) !" O. $' ()*+' M. -../ 0;': 1' &2 3"4"%56@ &' (8+' M -../ 0:'9O 1'. <=

(656)

(657) > 2?" $%&' ?2B" ?

(658) 4 &44

(659)

(660) C D ;#. 123456G89 KL2"

(661)

(662)

(663) !" #. $' ()*+' M. -../ 09'F 1' &2 3"4"%56@ &' (8+' M -../ 0O'9: 1'. <=

(664)

(665) > 2?" $%&' ?2B" ?

(666) 4 &44

(667)

(668) C D OH. 123456GG9 KL2"

(669)

(670)

(671) !" 9. $' ()*+' M. -../ FD'; 1' &2 3"4"%56@ &' (8+' M -../ FF'HO 1'. <=

(672)

(673) > 2?" $%&' ?2B" ?

(674) 4 &44

(675)

(676) C D O0. 123456GI9 KL2"

(677)

(678)

(679) !" DH. $' ()*+' M. -../ FE'F 1' &2 3"4"%56@ &' (8+' M -../ FF';H 1'. <=

(680)

(681) > 2?" $%&' ?2B" ?

(682) 4 &44

(683)

(684) C D OF. 123456GJ9 KL2"

(685)

(686)

(687) !" DD. $' ()*+' M. -../ FO'D 1' &2 3"4"%56@ &' (8+' M -../ F#'DF 1'. <=

(688)

(689) > 2?" $%&' ?2B" ?

(690) 4 &44

(691)

(692) C D O:. 123456GN9 KL2"

(693)

(694)

(695) !" D0. $' ()*+' M. -../ :D'E 1' &2 3"4"%56@ &' (8+' M -../ :D'FH 1'. <=

(696)

(697) > 2?" $%&' ?2B" ?

(698) 4 &44

(699)

(700) C D OE. 123456GP9 KL2"

(701)

(702)

(703) !" DF. (8+' M. -../ :0'0# 1' <=

(704)

(705) > 2?" A2B" ?

(706) 4 &44

(707)

(708) C D O;.

(709) 12345689

(710) !"#$"%&'($##$)" *+,- ./0.12 3 4 5$#6)$78& &")5$#6)$78&'9 ,:2 ;< < =>2 2222 ?0 1234568@ AB < < => <C, 4 D 4- 9 B , <, ,,2 ED , ,: B,, , FB , B, , D *F ,<12 *1 G , BH !"#$"%&'&IJ) *+K,- ./0L1 > M'($##$)"'*+,- ./0.12 *91 G , FB F M'&IJ) ,4 > M'($##$)" ,42 *1 G , FB F M'&IJ) ,4 > M'($##$)" DDN A,4 O '&IJ)- ; O M' ($##$)" A,4 O 9B <C,2 222222222222222222222222222222222222222222222222222 ?P 1234568Q R, DD ,:, 4, SBHS *M'&IJ) M'($##$)"1 SFS *,4 DD1 < FB, F < < B B,, ,DF2 222222222222222222222222222222222222222222222222222222222222222222222222 ?? 123456TU !"#$"%&'&IJ) *+K,- ./0L12 E,V D >B 2 AA O AB, A< =>2 + O .W 2 *X< AV,- .//?12 222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222 .WY 123456T8

(711) B2 R1 !"#$"%&'&IJ) *+K,- ./0L12 +1 !"#$"%&'($##$)"'*+,- ./0.12 A; O AB, ;2 + O .2 X< ;H4Z[<, *./L/12 2222222222222222222222222222222222222222222222222222 .WY 123456TT !"#$"%&'&IJ)2 GF AA D2 A;- BD \ ,2 E AA A; F42 3,V ] D V O 3,V ,C, H B 2 3,V ] B O 3,V ,C, H B ]D,2 22222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222 .WP 123456T^ !"#$"%&'($##$)"'9 DD2 GF AA D2 A;- BD ,2 E AA A; F42 3,V ] D V O 3,V ,C, H B 2 3,V ] B O 3,V ,C, H B ]D,2 22222222222222222222222222222222222222222222222222222222222222222222222222 .W? 123456T_ !"#$"%&'($##$)"'B 9, , 2 GF AA D2 A;- BD ,2 E AA A; F42 3,V ] D V O 3,V ,C, H B 2 3,V ] B O 3,V ,C, H B ]D,2 222222222222222222222222222222222222222222222222222222222222 .W/ 123456T` ;,F V B B, < => *AA12 GB, F< D < =F M'&IJ)'*+K,- ./0L122222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222222 ..W 123456T9 ;,F V > , B B, < => *AA12 GB, F< VBH D < =F BH M'($##$)"'*+,- ./0.1 B 9, , 2 2222222222222222 ... 123456T@ ;,F V > , B B, < => *AA12 GB, F< VBH D < =F BH M'($##$)"'*+,- ./0.1 D DD2 22222222222222222222222222222222 ....

(712) 1234567.

(713) . ! " # $% & ' &( & )*+,-./0123 2456*++* 78 9&: ;<=> )*+,-./0.123?0//06.* 78: ;< > ; @ AB A C& A. A A D! D%& A< A < !& A A E << <=

(714) FGHECI

(715) " # & %J DK B

(716) FGHECI 8 " # & %J L@ M= ! A " N & @ & !O' &@ & % P P & )*+,-./0.12 F%#"L: ;=; 7J F&> & & 8! @ ; < C& QB < A D! D%& QA < < !& QM < E Q; ;A ! < " D&& @!O$ @ !@% & @ & & % & )*+,-./0.12 F%# L: ;=; 7J F&> & & & 8! ; @ ;; C& BB A D! D%& BA < !& B= E . A . M.

(717) 13567898

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`J#%F#)-'.Z[][>/',",%%'.)#-/%#8$')-'.'$0:/#! 6%#!18'1%#!?A#::1,)1aA#"8'#).BCC\E3 '!-#+,+,b1,c%#)+'+,<,%-'.#85$+'#(%'!')-#%F,%-'#-121+#+'('!>/'1%#. -#$65$.(,!!/1/$:%#)+'",$(8'7,+'F#`')+#!+'"#%"1)1"/8-/%#.#!!/$1)+,/$#(,!1G;, +'+'!-#>/'')-%',!$#1,%'!(%,+/-,%'!+'"#$#%&'!"/8-12#+,!+,9%#!18?@94A4.BCCDE3 !$#1,%'!(,8,!+'(%,+/G;,!'",)"')-%#$)#!$#%:')!+,!'!-/0%1,!'%1,!",!-'1%,!+, 81-,%#8),%-'+,'!-#+,.,)+'!'"/8-12#.(%1)"1(#8$')-'.KLMNOPQRPSTUdRQQRXPLU?9,,)'.Z[\ZE. /$#'!(5"1'+'"#$#%;,,%1:1)0%1#+,,"'#),H#"*F1",4$'%1"#),3 "#$#%;,6%#)",+, H#"*F1",5/$#+#!'!(5"1'!+'"#$#%&'!$#%1)=,!$#1!"/8-12#+#!),$/)+,?J%'1-#!PMURef. BCCDE'/$#+#!(%1)"1(#1!'!(5"1'!+'"#$#%&'!",$'%"1#1!+,H#"*F1",g'!-'?I#88'!h i1$')'`'H'%'`hj)%1>/'`.BCCkE.+'21+,l!!/#!"#%#"-'%*!-1"#!+'#8-##+#(-#G;,.%0(1+, "%'!"1$')-,.",)2'%!;,#81$')-#%'F1"1')-''#8-#!-#7#!+'!,6%'212_)"1#'$",)+1G&'! #$61')-#1!#+2'%!#!?9%1::!PMURef.BCCmE3 4('!#%+#1$(,%-n)"1#!o"1,h'",)p$1"#.#1)-%,+/G;,+''!(5"1'!#8o"-,)'!,/ '7o-1"#!(#%#,"/8-12,(,+'('%$1-1%#1)2#!;,+'!!#!'!(5"1'!'$#$61')-'!#>/0-1",! 12.

(719) 13567898

(720) . !"#$%&'(#)*+,-..-/01' ('23"'4#56708+-9*:;;<= >?#6@A?08+-9*:;;B='"'C#67DE#F:;;:G CDH#:;;IGJ':;K;=$L23''# !(A"'("22'('M "6CDH#:;;I=$N!"(#"')*+,-..-/01 A"F"(A'(O2!P(A"2"'Q &"#("'(2"#("A2O(R' "''F?"'("P"'#"#(#'P(" '#6CDH#:;;IGSTQUT08+-9*:;KK=$ L(A"(OM"''#"'" 2('('!"MV"''"2 (A""#26)*+,-..-/01=$%((A""'#'' !")*+WXY/1881)*+,-..-/01(''#'!#R"$ @MM2#'(#"?'(A")18Z[0.-0\W ''M(("O!"('!"#A(O2#' F!]''"36J^"08+-9*K<_`=$ L(A""'""O"'!#R"&3!"(R&'"'A ")*+,-..-/01)*+WXY/1881"F"""?'FO' #2"'!2!"(Ca08+-9*6K<<;=$%#A'AM!' 'F'!#'("#''"F (#2"'('2?M'!''A# ''''''2#6@Q08+-9$K<<b=$> 'M('!R('#F('!"#'(" 12.

(721) 13567898

(722) . !"#"$%&'''&('&&)*'&+' &+&)',''&))+-.&+'&&')&)/'&)'/&0&)*'& 0'/&1 23'&++'+'45'&,&-.&5('&('& 6789:;;:<=>678?@A<>BB>1C+&D'&+&&&&+&&)*'&%E+ &')'%))/'-.&+''&&,'& &,'&)*'3('%&&+,&/&''++ ,&''1F,G&H&IG!""!$%)3)%''&&678 9:;;:<=>,'+&J'KL4M& )3+N)'O+4&$&+E+&)*'&P)&+ ','('P+&,&''&12)'%&&&+&'E+ )5'&&''&''++0,N45'%K&L&'% +E+*)'+'&,'%)'&%&)&/'/'&%0 &,/'/'+'&G'&)+''1)&&+) ,,''/&0&)*'3(')')-.&5('&I)4+#$ 5*'&I)4+&!Q$+&&)*'&D6>BRS=;:=T?3(''/$1 . 12.

(723) 13567898

(724) . . !"#!$%&!'"!"!("!)*+),-./)+0+-1*2*..342)0!+0!-/562)-!-789)20! :;<=>?@A?BCDEA@@AF?;!G**4-!HIJHK!4*!2*L5M-7*!-/9N0.)4*OM0PN40.!+-!Q04046)0OPN05-O MR0!Q*L5.)+0"!&!$N4+*!+0!(0S+-!,"!JT!4"!U!5"!VWUOVWI!XTTV"! Y!Z"!"#!%Z[Q!"!%"#!%(\!%"!]"#!($\^!%"!%"!^_`.)+)a09)*4!*b! c?@A?BCDC?<;d?eBC!G%)440-N/!HfVfK!04+!c?@A?BCDCghF;<<;!N.i-4.*0+!HIJV!Gj-205*+0K"! ]*N.40M!*b!Q.N/902-04!)*M*P_!,"!HT!5"!XfklXkJ!HIIT"! m\(!n"!j"#!](o(!Q"!$"#!'%\\!]!n"!$"!/*M09)*4!04+!2R0.029-.)a09)*4!*b! 4-p!L)2.*/09-MM)9-!M*2)!)4!9R-!n02)b)2!pR)9-!/R.)L5!:;<=>?@A?BCDEA@@AF?;!04+!2.*//O/5-2)-/! 0L5M)b)209)*4!)4!*9R-.!5-40-)+!/5-2)-/"!$*M-2NM0.!2*M*P_![*9-/!,"!f!5"!JXUlJXV!XTTf"! '&[qr%qOstuvwsrxyzv{|z}~vuyz{z{|z}~yzv{|zt vyz{z //-//L-49!*b!9R-!)49-.029)*4!`-9p--4!9R-!pR)9-!/R.)L5!cAA?F=@D=@;e=C<e;C!04+!9R-! -7*9)2!*.)-490M!/R.)L5!cAA?F=@DFAge=Ag<BC!)4!0!N.*5-04!-/9N0._!G(Z!(50)4K"!)*M*P_! 4,0/)*4/!,"!HX!5"!HfJHlHfUW!XTHT"! $"!/909/9)20!+-!n-/20!O!.0/)M"!'.04+-/!.-P)-/!-!N4)+0+-/!+0!b-+-.0*"! .0/M)0!5"!HWH!XTTf"! %Z[Q!"!%"!?<DA!(N22-//bNM!)49-./5-2)b)2!2.*//!*b!9p*!/5-2)-/!*b!L0.)4-! /R.)L5!n-40-N/!/9_M).*/9.)/!!n-40-N/!/-9)b-.N/"!042*N,-.!"Q"1!HW9R!44N0M!$--9)4P!*b!9R-! Z*.M+!$0.)2NM9N.-!(*2)-9_"!HIkU"!5"!JI"! %&$[[!j"#!$!"!Q"!'"#!%!]"!\"!\R-!)4,0/)*4!*b!b),-!0M)-4!/5-2)-/!)4!9R-! j-M90!+*!n0.40`0!4,).*4L-490M!n.*9-29)*4!.-0![*.9R-0/9-.4!.0a)M"!-,)/90!+-!)*M*P)0! \.*5)20M!,"!Wk!4"!J!5"!ITIlIXJ!XTHT"! 12.

(725) 13567898

(726) . !"#$%&'!()&*#+&#*,!- ./012345367879:;/00/<,+%"!=%> ,(%,'=%,?-*!@&A,B*%C'$'%(+!%)&'=,(&'-',=#)'(D )'E"!$F@!*"A'+@'+*!)%&,$$'&,$!+'!$,+#$%*+!$!DFGHI"JIHJKJIHLIMMJ NO POQRSR(&*!=#TU!% %(V$'),)@!*-!@W&*'+%),+!$!D'%#)&*%$')GB*%)'$P!$#@,>OX"'+!),@&*%&%@,(&!)=, =%=!)Y'!$XD'+!)'!=,Q%(,'*!>ZRQ"[LN\]H]][ ^SQRS_O`BQ B)",+',)@%*'(%),EX&'+%)F+*'"&!DW('+%),($%)+!)&%)=, P,(,C#,$%B!$,&'(=,$()&'&#&!=,+,%(!D*%-'%P,(,C#,$%Gab(H"L]K]bIMML ORc )"W+',),EX&'+%)=,+%@%*d,)@%*'(A!) <e3453678;141f1;R%Y*'+'#)HL]\,./0123453678g5445;3/B!!(,H]JH?(!)%@Y',(&,) ,)&#%*'(!,@%*'(A!=!(!*=,)&,=!B*%)'$B!$,&'@OW+('+!N+',(&h-'+!=! O%@%(=%*WGHM(H"IM]NIIIIMMI iO__NZO_ZBOO iiiOiZOc_OQOA,"*,),(+,!-&A, "%+'-'+jA'&,$,D)A*'@"<./0123453678g5445;3/B!!(,H]JH?'(&A,j'$='(OA%'$%(= Q!#*(%$!-A,$$-')A,),%*+AGIb(a"HH\LNHH]IIMML PNQSZS SNkZS !"#$%&'!(,(,&'+ &*#+&#*,!- %+'-'+iA'&,A*'@"<./0123453678g5445;3/?-*!@,E'+!&! %(%@%> '+*!)%&,$$'&,P%*'%&'!(%*'(,B'!&,+A(!$!DFGb"aL[Ka\aIMM[ i_N_ZZ_OQBc k*,),(+,!-%+'-'+jA'&,)A*'@"./0123453678g5445;3/<B!!(,H]JH?'(&A,!#&A,*( #$-!-,E'+!l#%&'+(G%)'!()Gb(H"HJ]KHaIIMHH 12.

(727) 3456789

(728) 34474. 7 74 447 84 8 !"#$%&%'() *+8, 4-

(729) ./01 !"#!$%&2#""#)! *+. -

(730) ./

(731) 1. 3456789:;<;. 34=437>=?@4AB:; :C7:D>CE7?34; F:G; 34=437>=>G; G>H84?G; G>38CFI=?:G;F>;JKLMNOPQRSTSUVWKLLKTXY8=Z>C=:4F[;<\]^_;>; JKLMNOPQORST`QPPQWOKTXY::C>[;<\]<_;. 4abc;Ydefghd;ib;jgedbh;k;jgedbh;X4l;Yl_; Fdmnbab;3ghob;Ggdebh;ib;jgedbh;k;jgedbh;XFl;3l;Gl_; Fdmnbab;Ggfebned;k;Ggfebned;XFl_; 4aaphhgm;5gmqbh;5nmrbneg;k;5nmrbneg;X5l;5l_; staung;4vewang;ib;jgednh;sebneb;k;sebneb;Xsl;4l;jl_;. xy74z + 9 49 {y4 *|

(732) 1. ;. 12.

(733) 3456789

(734) 34474. 7 74 447 84 8 !"#$%&%'() *+8, 4-

(735) ./01 !"#!$%&2#""#)! *+. -

(736) ./