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290

Pigozzoi Almeida-Silva, Griswold & Brescovit gen. nov.

Etymology. The generic name masculine and is in honor to Raphael Pigozzo, Lina's

fiancé.

Type-species. Pigozzoi pifozonho Almeida-Silva, Griswold & Brescovit sp. nov.

Diagnosis. Species of the genus Pigozzoi gen. nov. differ from all other

Macrobuninae by having a retrolateral subtegular apophysis and by having the RTA fused to the iRTA.

Description. Total length: 2.57-3.45. Anterior and posterior eye rows straight. AME

as large as or larger than ALE, PME and PLE. All indirect eyes with canoe shaped tapetum, prolaterally displaced in PME. Coloration yellowish to pale brown, with or without chevron marks at posterior area of abdomen; legs with alternating dark rings. Chilum absent. Clypeal hood absent. Chelicerae not geniculate. Promargin with four teeth, retromargin with one or two denticles and four to five teeth. Sternum pentagonal, projected between legs IV. Leg spination strong, with long spines arising from prominent bases (Fig. 138B). Trocanther distal margin straight.

Calamistrum short. Cribellum present, entire, wider than long. Tracheal spiracle with 1/3 the size of distance between the ALS outer margin. Median tracheae branched.

PLS larger than PMS but smaller than ALS. Male with cymbium subtriangular, with a median basal apophysis, curved to the prolateral side, and a prolateral knob that interacts with the PTA. MA rectangular with prolaterally curved apex (Fig. 137A, E, F-G). Distal tegular process membranous and short (Fig. 137F-H). TA1 reduced (Fig. 137F-H). Embolus short and thick, rising on prolateral apical side of tegulum.

Subtegulum bearing a prolateral apophysis. PTA present and interacting with a

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291 prolateral knob on the cymbium (Fig. 137C). VTA absent. RTA and iRTA fused, triangular (Fig. 137E). DTA globose, with a saw striae on the retrolateral side and pointing to the prolateral side (Fig. 137B, C, E). Female epigynum with rounded to oval spermathecae and sinuous copulatory ducts (Fig. 139D). Median an lateral lobes distinct. Middle-field with posterior plate. Middle-fied central area subrectangular with expansions to the copulatory opening area.

Distribution. Chile and Argentina.

Composition. Monotypic.

Undescribed species. Three.

Pigozzoi pifozonho Almeida-Silva, Griswold & Brescovit sp. nov.

Type-material. Holotype male, paratype female from 20km E from Potrero Grande,

El Relvo, Curicó, Región de Maule, Chile, 26.III.2004, N. Dumbeyl & J. E. Barriga- Tuñol coll., deposited at CAS (CASENT 9032831); Paratype male ande female with the same data, deposited at IBSP.

Etymology.The specific name is a ramdom combination of letters.

Diagnosis. Pigozzoi pifozonho differs from other species by the pointed and small

embolic base apophysis (Fig. 137D, F-H).

Description. Male (Holotype). Total length 3.25. Carapace piriform, length 1.45,

width 1.28, height 0.50. Fovea short slightly deep. Eye rows as in generic description. Eye diameter and interdistances: AME 0.08; ALE 0.10; PME 0.09; PLE 0.10; AME-AME 0.04; AME-ALE 0.03; ALE-PLE 0.03; AME-PME 0.06; PME-PLE

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292 0.06; PME-PME 0.10. Tapetum as in generic description. Coloration as in generic description except abdomen grayish with three pairs of pale spots on its median dorsal line. Clypeus 0.10 high. Clypeal hood absent. Chelicerae: fang 0.28; paturon 0.35, not geniculate. Promargin with four teeth, retromargin with two denticles and five teeth (Fig. 138C). Labium: 0.14 long, 0.29 wide, straight, slightly rebordered.

Leg measurements. I: 1.38 femur/ 0.53 patella/ 1.48 tibia/ 1.13 metatarsus/ 0.50 tarsus/5.00 total; II: 1.22/ 0.54/ 1.28/ 0.87/ 0.63/ 4.55; III: 1.12/ 0.44/ 0.92/ 0.96/

0.36/ 3.80; IV: 1.30/ 0.44/ 1.22/ 1.16/ 0.46/ 4.58. Leg formula: 1423. Left legs and palp spination: Femur: IV: 0-0-1pap; Tibia: I-II: v 2-2-2-1p; III: v 0-2-0-2ap; IV: 0- 1-2ap, d 1sap; p 0-1; Metatarsus: I: v 2-2-2-0; II: v 2-2-2-1map; III: v 2-2-1map, p 0-1-0-2; IV: v 0-2-0-1map, r 1sap; p 1-0-1sap. Tarsal organ flat and small (Fig.

138D). Trichobothrium hood longitudinally striate (Fig. 138F). Third claw with four acessory teeth (Fig. 138F). Spinnerets with reduced number of spigots (Fig. 140).

Cribelum modified in colulus (Fig. 140A). ALS with one MAP a nubbin and four Pi spigots (Fig. 140B). PMS with one mAP and no other spigots (Fig. 140C). PLS with MS reduced to a nubbin an flanked by a nubbin and two AC spigots (Fig. 140D).

Male palp as in generic description except for embolic base apophysis shape (Fig.

137D, F-H), median apophysis apex concave (Fig. 137G) and distal tegular process shape (Fig. 137F-H).

Female (Paratype). Total length 2.88. Carapace as in male, length 1.34, width 1.13, height 0.70. Fovea and eye rows as in male. Eye diameters and interdistances: AME 0.09; ALE 0.09; PME 0.08; PLE 0.10; AME-AME 0.04; AME-ALE 0.02; ALE-PLE 0.03; AME-PME 0.07; PME-PLE 0.06; PME-PME 0.10. Tapetum as in generic description. Coloration as in male, except abdomen with chevron marks at the end.

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293 Fig. 137: Pigozzoi pifozonho sp. nov., ♂, left palp. A. Bulb, apical view; B. DTA tip; C. Palp, dorsal view; D. Bulb, prolateral virw; E. Palp, retrolateral view; F. Bulb, ventral view; G.

Palp, ventro apical view; H. Bulb apex.

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294 Fig. 138: Pigozzoi pifozonho gen. nov.: A. ♂, right leg IV; B. ♂, right tibia I; C. ♂, left chelicerae, retromargin; D. Right leg IV, tarsal orga; E. clawEyes, ♂; F. Right leg IV, metatarsal trichobothrium.

Clypeus 0.07 high.Clypeal hood absent. Chelicerae: fang 0.23; paturon 0.40, not geniculate. Promargin with four teeth, retromargin with one denticle and four teeth.

Leg measurements. I: 1.10 femur/ 0.50 patella/ 1.24 tibia/ 0.89 metatarsus/ 0.41 tarsus/ 4.14 total; II: 1.00/ 0.44/ 0.96/ 0.81/ 0.38/ 3.59; III: 0.85/ 0.40/ 0.77/ 0.74/

0.37/ 3.13; IV: 1.24/ 0.45/ 1.06/ 1.03/ 0.42/ 4.20. Leg formula: 4123. Left legs and palp spination: Femur: I: 0-0-1p; Tibia: I-II: v 2-2-2-1p; III: v 0-1r-0; IV: v 0-1r-

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295 1rap; Metatarsus: I: v 2-2-2; II: v 2-2-2-1msap; III: v 2-2-0-1r, r 1sap, p 0-1-0-1sap;

IV: v 1map, r 0-1-0-1sap, p 1sap. Cribellum entire (Fig. 141A). ALS with two MAP and five Pi (Fig. 141). PMS with one mAP and one CY (Fig. 141C). PLS with one Cy, two AC and one MS flanked by a nubbin (Fig. 141D). Epigynum middle-fied and lateral lobes distinct. Middle-field shape as in Fig. 139A-B. Middle-field with posterior area not dorsally projected or forming a plate (Fig. 139C). Spermathecae oval, copulatory ducts sinuous (Fig. 139D).

Variation. Length (10♂): Total: 2.68-3.28; Carapace: 1.25-1.56; Femur I: 1.10-1.30.

Length (10♀): Total: 2.90-3.58; Carapace: 1.19-1.47; Femur I: 0.81-1.35.

Additional material examined. CHILE. Región VII de Maule, Curicó, El Relvo,

20km E from Potrero Grande, 50 ♂, 47♀, 26.III.2004, N. Dumbeyl & J. E. Barriga- Tuñol coll., CAS (CASENT 9032831; 9021225); 2♀, 2♂, 18.VI.2004, CAS (CASENT 9020937); 6♀, 2♂, 13.VI.2004, CAS (CASENT 9032179).

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296 Fig. 139: Pigozzoi pifozonho, ♀. A. Epigynum ventral view; B. Epigynum, ventro anterior view; C. Epigynum posterior view; D. Vulva, dorsal view.

Fig. 140: Pigozzoi pifozonho sp. nov., ♂, spinnerets. A. Overview; B. ALS; C. PMS overview; D. PLS.

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297 Fig. 141: Pigozzoi pifozonho sp. nov., ♀, spinnerets A. Overview; B. ALS; C. PMS overview; D. PLS.

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298

Samadhia Almeida-Silva, Griswold & Brescovit gen. nov.

Type-species. Samadhia kellyfujiharae sp. nov.

Diagnosis. Males of Samadhia differ from all Macrobuninae by the presence of

tibial crack. Samadhia resembles Naevius, Emmenomma, Anisacate, Yupanquia and Ijoubetumsam by the presence of a gland in the palpal tibia which discharges

through the DTA; differ from Emmenomma and Naevius by the DTA fixed to the tibia; differs from Ijoubetumsam by the presence of AME; differ from Anisacate by the long and oblique cheliceral groove and entire cribellum; differ from Yupanquia by the narrow and pointed DTA.

Etymology. The generic name is femine and is an honor to Samadhi Dance school in

São Paulo.

Description. Total length: 2.90-5.56. Anterior and posterior eye rows straight. AME

smaller than ALE, PME and PLE. All indirect eyes with canoe shaped tapetum median in PME. Coloration yellowish to brown, with or without chevron marks at posterior area of abdomen, legs uniform. Chilum absent. Clypeal hood absent.

Chelicerae not geniculate. Promargin with three teeth, retromargin with three or four denticles and four to six teeth. Sternum pentagonal, projected between legs IV.

Trocanther distal margin straight. Calamistrum short. Cribellum present, entire, wider than long. Tracheal spiracle with 1/3 the size of distance between the ALS outer margin. Median tracheae branched. PLS larger than PMS but smaller than ALS. Male with cymbium subtriangular, with a prolateral knob (Fig. 142D). MA, large, with several complex projections and located in the middle of the tegulum (Fig. 142H). Distal tegular process and conductor absents. TA1 rectangular, curved,

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299 and bordering the embolus (Fig. 142E-G). Tegular rim short and located on the retrolateral side of the tegulum (Fig. 142G) (Fig. 142F). Embolus short, arising on prolateral apical side of tegulum and bearing a hyaline expansion on the tip (Fig.

142E, G). Subtegulum prolateral. PTA present. VTA absent. RTA reduced to knobs (Fig. 142B-C). iRTA sub triangular (Fig. 142B-C). DTA long or short, with one branch and bearing an opening on its tip (Fig. 142A). Tibia globose, with a gland that discharges through an onpening on DTA (Fig. 142A-D, I). Female epigynum with rounded to oval spermathecae and sinuous copulatory ducts. Median and lateral lobes fused or distinct with median lobe slightly projected over lateral lobes (Fig.

146A). Middle-field posterior region dorsally projected and forming a plate (Fig.

146B) or not.

Distribution. Chile and Argentina.

Composition. Monotypic.

Undescribed species. Two.

Samadhia kellyfujiharae Almeida-Silva, Griswold & Brescovit sp. nov.

Types. Holotype male, paratype female from Palmas de Oca, Valparaíso, Chile, 31-

VIII.1984, R. Calderón coll., deposited in AMNH (AMNH); Paratypes male and female with same data, deposited in IBSP.

Diagnosis. Samadhia kellyfujiharae differs from other species of the genus by the

shape of its MA with a broad rugose projection (Fig. 142H); and by the shape, length and girth of the DTA (Fig. 142D) in the male palp and by the posterior area of the epigynum dorsally projected and forming a square plate (Fig. 146B).

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300 Etymology. The specific name is an honor the belly dancer and teacher Kelly

Fujihara.

Description. Male (Holotype). Total length 4.08. Carapace piriform, length 2.12,

width 1.53, height 0.96. Fovea short and slightly deep. Anterior eye row slightly procurved, posterior straight (Fig. 143A). Eye diameters and interdistances: AME 0.06; ALE 0.10; PME 0.09; PLE 0.10; AME-AME 0.03; AME-ALE 0.04; ALE-PLE 0.04; AME-PME 0.06; PME-PLE 0.09; PME-PME 0.10. Tapetum median.

Coloration: carapace brown, with faint pigmentation on radial muscular impression and delimitation of the cephalic area; legs, chelicerae, endites, and labium brown;

sternum grayish; abdomen with paired spots around the cardiac area, forming chevron marks on its end. Clypeus 0.14 high. Clypeal hood absent. Chelicerae: fang 0.51; paturon 0.88, not geniculate. Promargin with three teeth , retromargin with four denticles and four to five teeth. Labium: 0.29 long, 0.29, notched, not rebordered. Leg measurements. I: 2.03 femur/ 0.70 patella/ 2.10 tibia/ 1.78 metatarsus/ 1.00 tarsus/ 7.61 total; II: 1.64/ 0.62/ 1.56/ 1.42/ 0.75/ 5.99; III: 1.47/

0.63/ 1.24/ 1.44/ 0.63/ 5.42; IV: 2.10/ 0.69/ 1.86/ 2.24/ 0.86/ 7.74. Left legs and palp spinulation: Femur: I-II: d 0-1-1p; III: d 0-1-3; IV: d 0-1-2; Patella: III-IV: d 0-0-1;

Tibia: I: v 2-2-0-2ap, p 1v-1d; II: v 1m-1m-2ap, p 0-1d-1r-1d-0; III: v 2-2-2ap, d 1- 0-0-1-0, p and r 0-1-0-1-0; IV: v 2(1r-1p)-1p-2ap, d 1-0-0-1-0, p and r 0-1-0-1-0.

Colulus rectangular (Fig. 144A). ALS with one MAP, one nubbin and nine Pi (Fig.

144B). PMS with one mAP and five AC (Fig. 144C). PLS with MS and flankers reduced to nubbins and ten AC spigots (Fig. 144D). Tibial apophysis and palpal sclerites as in diagnosis and generic description (Fig. 142).

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301 Fig. 142: Samadhia kellyfujiharae, ♂, right palp. A. DTA tip; B. Tibia, retrolateral view; C.

Tibia, ventral view; D. Tibia, dorsal view; E. Bulb apex; F. Bulb, ventral view; G. Bulb, retrolateral apical view; H. Median apophysis, ventral view; I. Tibia, prolateral view, showing tibial gland (light microscopy image, scale: 0.25mm).

Female: Total length 5.31. Carapace as in male, length 2.05, width 1.43, height 0.72.

Fovea short and shallow. Eye rows straight. Eye diameters and interdistances: AME 0.06; ALE 0.11; PME 0.09; PLE 0.11; AME-AME 0.04; AME-ALE 0.04; ALE-PLE 0.06; AME-PME 0.06; PME-PLE 0.10; PME-PME 0.11. Tapetum and coloration as in male. Clypeus 0.11 high. Clypeal hood absent. Chelicerae: fang 0.52; paturon

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302 0.89, geniculate. Promargin with three teeth, retromargin with three denticles and six teeth (Fig. 143B). Labium: 0.28 long, 0.31 wide, notched, not rebordered.

Fig. 143: Samadhia kellyfujiharae sp. nov. : A. Carapace, eyes, ♂; B. Left chelicerae, ♀, retromargin; C. Right leg IV, ♂; D. Right leg IV, ♀, tarsal trichobothrium.

Leg measurements. I: 1.60 femur/ 0.63 patella/ 1.49 tibia/ 1.20 metatarsus/ 0.71 tarsus/ 5.64/ total; II: 1.32/ 0.59/ 1.13/ 0.98/ 0.59/ 4.60; III: 1.15/ 0.52/ 0.90/ 1.02/

0.52/ 4.12; IV: 1.50/ 0.63/ 1.37/ 1.48/ 0.63/ 4.61. Left legs and palp spination.

Femur: I-II: d 0-1-1p; III: d 0-1-2; IV: d 0-1-0; Tibia: I: v 2-2-0, p 0-1-0; II: v 1r-1r;

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303 III: v 2-1p-2ap; r and p 0-1-0-1-0, d 1-0-0-1-0; IV: v 1p-2-2ap, r and p 0-1-0-1-0, d 1-0-0-1-0; Metatarsus: I: v 2-2-1map, p 1ap, r 1ap; II: v 2-2-1map, r 1sap, p 0-1- 2sap; III and IV: v 2-2-1map, r 1sap, d 2-2-2sap, p 0-1-1sap.

Fig. 144: Samadhia kellyfujiharae sp. nov., ♂ spinnerets. A. Overview; B. ALS; C. PMS; D.

PLS.

Cribellum entire and rectangular (Fig. 145A). ALS with two MAP and 11 Pi (Fig.

145B). PMS with two Cy, one mAP and eight AC spigots (Fig. 145C). PLS with one Cy, MS and one of the AC-like flankers sharing the same base and the other flanker separate (Fig. 145D). PLS with several AC spigots (Fig. 145D). Middle-field slightly

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304 projected over lateral lobes and both not fused as a plate (Fig. 146A). Posterior area of the epigynum dorsally projected and forming a plate (Fig. 146B). Copulatory ducts sinuous (Fig. 146C-D).

Fig. 145: Samadhia kellyfujiharae, ♀, spinnerets. A. Overview; B. Left ALS; C. Right PMS;

D. Left PLS.

Variation. Length (10♂): Total: 3.75-5.56; Carapace:1.78-2.01; Femur I: 1.44-1.58.

Length (10♀): Total: 2.90-3.58; Carapace: 1.19-1.47; Femur I: 0.81-1.35.

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305 Additional material examined. CHILE. Valparaíso, Palma de Ocoa, 38♂, 7♀, 12- 31.VIII.1984, R. Calderón coll., AMNH (12 vials).

Fig. 146: Samadhia kellyfujiharae sp. nov., ♀, genitalia. A. Epigynum, ventral view; B. Ditto, posterior view; C. Vulva, dorsal view; D. Vulva, left CD and spermathecae.

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306

Segoko Almeida-Silva, Griswold & Brescovit gen. nov.

Type-species. Segoko chueneorum Almeida-Silva, Griswold & Brescovit sp. nov.

Diagnosis. Species of Segoko gen. nov. differ from all other Macrobuninae except Neoporteria, Anisacate, Pseudauximus, Pigozzoi and Imbumbulu gen. nov by the

AME larger than the PME (Fig. 147E); differs from Anisacate and Neoporteria by the lack of median apophysis; differs from Pseudauximus and Imbumbulu by the shape of the TA2 sub rectangular (Fig. 148H) and DTA subtriangular with one retrolateral branch (Fig. 148A). Female epigynum resembles those of Pseudauximus and Imbumbulu in having a plate surrounding the mid-field but differs from Pseudauximus by the membranous middle-field and from Imbumbulu by the presence

of a rounded membranous median projection on the middle-field (Fig. 149A-C).

Etymology. The generic name is neutral, from the Northern Sotho word for spider.

Description. Total length: 4.15 - 5.30 Anterior and posterior eye rows procurved.

AME as large or larger than PME, PLE and ALE. (Fig. 147E). Both AME surrounded by a strong and contiguous dark circle. All indirect eyes with canoe- shaped tapetum inner displaced. Coloration yellowish brown, legs with alternating dark rings on the ventral face of the femur. Chilum entire or absent. Clypeal hood present. Chelicerae geniculate; promarging with five teeth, retromargin with two denticles and three teeth. Labium slightly rebordered or not and notched. Sternum pentagonal, longer than wide and projected between legs IV. Trochanther unnotched.

Calamistrum reduced to the base of metatarsus IV. Cribellum entire, wider than long, with width equal to 1/2 of the distance of the outer margin of ALS. PLS smaller than ALS but much larger than PMS. Male palp with cymbium oval, without

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307 apophysis but with a prolateral knob. Median apophysis absent. Distal tegular process membranous and embracing tip of emboluS (Fig. 148F). Embolus short, arising apically on the tegulum with a apophysis in the base and a constriction at tip.

TA1 and TA2 presents and well developed (Fig. 148F). Tegulum elongated with all sclerites concentrated on apical region (Fig. 148I). Subtegulum prolateral, with a depression (Fig. 148D, I). DTA triangular with a branch strongly projected to retrolateral side (Fig. 148A, C). RTA subrectangular, convex with a knob attached and delimiting a depressed area (Fig. 148C). iRTA subtriangular with a roundish tip.VTA and PTA present (Fig. 148B). Female epigynum weakly sclerotized and composed of a median membrabous area with a rounded projection surrounded by a sclerotized area (Fig. 149A-C). Spermathecae oval, longer than wide (Fig. 149D).

Copulatory ducts straight (Fig. 149D). Primary pores scattered around the dorsal area (Fig. 149D).

Composition. Monotypic.

Distribution. South Africa: Limpopo.

Undescribed species. One.

Segoko chueneorum Almeida-Silva, Griswold and Brescovit sp. nov.

Type-material. Holotype ♂ from Marbel Hall, South Africa, 25.V.1999, P. Stephen

col., Deposited in PPRI (AcAT 99/244). Paratype ♀ with the same locality, collection data and in the same vial. Paratypes: 2 ♀ 1 ♂, with the same data from holotype, deposited in IBSP.

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308 Etymology. Speciecific name is a honor for our friends Thomas and Kgabi Chuene

and their family, from Polokwane, Limpopo

Diagnosis. Segoko chueneorum differs from other species of the genus by the

appearance of its TA2 which is smooth and does not have denticles (Fig. 148F, H).

Females of other species are unknown.

Description. Male (Holotype). Total length 4.05. Carapace piriform, length 1.95,

width 1.50, height 0.86. Thoracic fovea deep and short. Anterior and posterior eye rows procurved. Eye diameters and interdistances: AME 0.13; ALE 0.1; PME 0.08;

PLE 0.10; AME-AME 0.03; AME-ALE 0.04; ALE-PLE 0.03; AME-PME 0.08;

PME-PLE 0.08; PME-PME 0.09.Tapetum prolaterally displaced on PME. Coloration as in generic description but also includes: abdomen grayish with indistinct pattern;

carapace with pigmentation over radial muscular impressions and delimitation of the cephalic area. Clypeus 0.07 high. Clypeal hood present. Chelicerae: fang 0.34;

paturon 0.55 paturon, geniculate. Promargin with five teeth, retromargin with two denticles and three teeth. Leg measurements: I: 1.66 femur/ 0.72 patella/ 1.78 tibia/

1.75 metatarsus/ 1.03 tarsus/ 6.94 total; II: 1.44/ 0.6/ 1.36/ 1.41/ 0.84/ 5.65; III:

1.38/ 0.55/ 1.05/ 1.33/ 0.70/ 5.00; IV: 1.87/ 0.66/ 1.63/ 1.84/ 0.84/ 5.83. Leg formula: 1423. Left legs and palp spination: Femur: I: d 0-1-2(1p-1m); II-IV: d 0-1- 3; Patella : II-IV: d 0-0-1; Metatarsus: I: v 2-2-2ap, r 1sap, p 0-1-1sap; II: v 2-2-2ap, r 1sap, p 0-1-1sap, d 0-1-1sap; III-IV: v 1p-2-2ap, r and p 0-1-0-1-0, d 1-0-0-1-0;

Metatarsus: I: v 2-2-0, r and p 0-1-0-2ap; II: v 2-2-1map, r 0-1d-1-1d-2ap, p 1d-1d- 2ap; III-IV: v 2-2-1map, d d 2-2-2 or 2-3-2, p 1ap, r 1ap; Tarsus III-IV: six to nine ventral spines. Cribelum reduced into a colulus (Fig. 151A). ALS and PLS similar in size, PMS smaller tha both (Fig. 151A). ALS with one MAP one nubbin and seven Pi

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309 (Fig. 151B). PMS with one mAP and four AC (Fig. 151C). PLS with MS reduced to a nubbin and some AC spigots (Fig. 151D).

Fig. 147: Segoko chueneorum sp. nov. A.Tarsal organ, right leg I, ♀; B. Right leg I, ♀, metatarsal trichobothrium; C. Right leg I, ♀, claw; D. Right chelicerae, ♂, retromargin; E.

Carapace, ♂, eyes; F. Right leg IV, ♀, calamistrum.

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310 Fig. 148: Segoko chueneorum gen. nov., ♂, left palp: A. Tibia, dorsal view; B. Tibia prolateral view; C. Tibia, retrolateral view; D. Tibia, ventral view; E. iRTA, retrolateral view;

F. Bulb, apical view; G.Bulb, retrolateral view; H. Bulb, prolateral view; I. Bulb, ventral view.

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311 Female (Paratype, PPRI). Total length 4.16. Carapace as in male, length 1.76, width 1.24, height 0.58. Fovea and eye rows as in male. Eye diameter and interdistances:

AME 0.09; ALE 0.10; PME 0.08; PLE 0.08; AME-AME 0.03; AME-ALE 0.03;

ALE-PLE 0.06; AME-PME 0.06; PME-PLE 0.08; PME-PME 0.11. Clypeus 0.05 high. Tapetum position as in male. Coloration as in male, except posterior area of the abdoment with indistinct chevron marks. Chelicerae fang 0.36; paturon 0.62 paturon. Teeth and denticles as in male. Labium: 0.21 long, 0.26 wide, notched, slightly rebordered. Leg measurements. I: 1.40 femur/ 0.62 patella/ 1.28 tibia/ 1.00 metatarsus/ 0.76 tarsus; II: 1.00/ 0.34/ 0.75/ 0.94/ 0.50; III: 1.13/ 0.53/1.00/ 0.82/

0.60; IV: 1.38/ 0.50/ 1.22/ 1.30/ 0.70. Left legs and palp spination: Femur: I-II: d 0- 1-1p; III-IV: 0-1-3; Patella III-IV: d 0-0-1; Tibia: I: v 2-2-2-0; II: v 1-1-0, d 0-1-1;

III: v 1-1-2ap, p and r 0-1-0-1-0, d 1-0-0-1-0; IV: v 2(1r-1p)-0-2ap, p and r 0-1-0-1- 0, d 1-0-0-1-0; Metatarsus: I: v 2-2-2ap; II: v 2-2-1map, p 1ap, d 0-1-1, r 1ap; III-IV:

v 2-2-1map, d 2-2-2sap, r 1sap, p 0-1-0-1sap. Tarsus as in male. Tarsal organ flat (Fig. 147A). Trichobothrium hood longitudinally striate (Fig. 147B).Third claw with four acessory teeth (Fig. 147C). Calamistrum reduced to the base of metatarsus IV (Fig. 147). Cribellar plate rectangular with spigots field anterior margin concave (Fig. 150A-B). ALS with two MAP and four to six Pi spigots (Fig. 150C). PMS with one mAP, one PC, two Cy and five AC spigots (Fig. 150D-E). PLS with one Cy and ons MS without flankers (Fig. 150F). Epigynum as in generic description (Fig. 149).

Variation. (2♂): Total: 4.05-4.15; Carapace: 1.95-2.01; Femur I: 1.60-1.66. Length (6♀): Total: 4.16-5.44; Carapace: 2.06-2.40; Femur I: 1.40-2.05.

Additional material examined. SOUTH AFRICA: Limpopo, Lajuma, Polokwane (23º 03' S; 29º 45' E): 1 ♂ 5 ♀, 15.VI.1997 (AcAT98/1); 1 ♂, 29.VI.1997

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312 (AcAT98/2); 2 ♂, 17.V.1997 (AcAT98/3); 2♂ 2♀, 13.VII.1997 (AcAT98/4); 7♂, 01.VI.1997, (AcAT98/5), all collected by M. vd Merwe; Limpopo, Nature Reserve (23º 9' S; 29º 47' E), ♀, 23.VI.2005, T. Khoza & M. Modiba coll, deposited in PPRI (AcAT2008/1392).

Fig. 149: Segoko chueneorum sp. nov., ♀, genitalia. A. Epigynum, ventral view; B.

Epigynum, ventro posterior view; C. Left copulatory opening; D. Vulva, dorsal view.

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313 Fig. 150: Segoko chueneorum sp. nov., ♀, spinnerets. A. Overview; B. Cribellum; C. Right ALS; D. PMS overview; E. Left PMS; F. Right PLS.

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314 Fig. 151: Segoko chueneorum sp. nov., ♂, spinnerets. A. Overview; B. Left ALS; C. Left PMS; D. Right PLS.

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