UNIVERSIDADE FEDERAL DA PARAÍBA
CENTRO DE CIÊNCIAS AGRÁRIASPROGRAMA DE PÓS-GRADUAÇÃO EM AGRONOMIA
C
OMPOSIÇÃO
F
LORÍSTICA,
F
ITOSSOCIOLOGIA E
I
NFLUÊNCIA
DOS SOLOS NA ESTRUTURA DA VEGETAÇÃO EM UMA ÁREA
DE
C
AATINGA NO
S
EMI-ÁRIDO
P
ARAIBANO
Lúcio Valério Coutinho de Araújo
Areia – Paraíba
Brasil
UNIVERSIDADE FEDERAL DA PARAÍBA
CENTRO DE CIÊNCIAS AGRÁRIASPROGRAMA DE PÓS-GRADUAÇÃO EM AGRONOMIA
COMPOSIÇÃO FLORÍSTICA, FITOSSOCIOLOGIA E INFLUÊNCIA
DOS
S
OLOS NA
E
STRUTURA DA
V
EGETAÇÃO EM UMA
Á
REA
DE CAATINGA NO SEMI-ÁRIDO PARAIBANO
Lúcio Valério Coutinho de Araújo
Orientador:
Prof. Jacob Silva Souto, D.Sc.
Tese apresentada à Universidade Federal da Paraíba, Centro de Ciências Agrárias, como parte das exigências à obtenção do Título de Doutor em Agronomia, Área de Concentração Ecologia Vegetal e Meio Ambiente.
Areia – Paraíba
Brasil
COMPOSIÇÃO FLORÍSTICA, FITOSSOCIOLOGIA E INFLUÊNCIA DOS
SOLOS NA ESTRUTURA DA VEGETAÇÃO EM UMA ÁREA DE CAATINGA
NO SEMI-ÁRIDO PARAIBANO
Tese aprovada pela Comissão em 15 de fevereiro de 2007.
B
ANCAE
XAMINADORAProf. JACOB SILVA SOUTO, D. Sc.
Unidade Acadêmica de Engenharia Florestal – UFCG
Prof. EVERARDO V. S. B. SAMPAIO, Ph. D. Departamento de Energia Nuclear – UFPE
Prof. RÔMULO SIMÕES CEZAR MENEZES, Ph. D. Departamento de Energia Nuclear – UFPE
Prof. DJAIL SANTOS, Ph. D.
Departamento de Solos e Engenharia Rural – CCA/UFPB
Prof. LEONARDO PESSOA FELIX, D. Sc. Departamento de Fitotecnia – CCA/UFPB
! " # $%
# & ' ( " & ) *
+ " ,
-. + ' # & ) . /0(& *
1 2 &
"- '
' " ' 3 ( " &
' ) * ( " & '
4 ) *
5 " '
$% & ' ( " & 6 " 6 " ) 24
" ,!!! "
7 & ' . ' # 86
9 : ( " .% ).
$ ,!!; + #
1 $% 8 # < # 2 :
( " & 7 # 1 ) *
!
"
# $
% %
& ' (
% % )
$
* +
,- . / 0
1 , 0 2 "
3 "
+ ' 3 7 ' $ ' $% ' '
" " >
( " & ' 4 ' $% '
' # ? (
7 # & >
+ @ . " . ' $% ' $
' " " ' >
.0 A + ' % %
' >
7 # 1 B(& *) C ' ' $%
D )# $% >
* B C ' ' ' % '
% >
# # A" ' " "7 '
" B *C>
+ 6 B& 1C # " '
% 1 ' ' 3 $%
' ? >
# " 6 ? ' 1" 3 $%
- >
1 & 1 ' 2 - . "
/ 2 ? % $ ' "
>
8E # : 8E : ? ' '
$%
'-# % >
? ' $%
6A. + &A4(/ . "
6A. + * 6 . 3
/ .(2F 3
*. / 3
AE /F+(GHF 4 / 6 I F * # I , & # # = , & = , , & # ; . # ! " $% # ,
, / & /JE A . *A*6AF4/K&A . I !" !" # $ % & '( # ,! / .(2F ,! *. / , AE /F+(GHF ,, , 2 /A 6 2L F+F. , , $% 1 ,
, , # ' $% ,I ; / .(6 +F. +A. (..HF , FE 6(.M . ;I I / & /JE A . *A*6AF4/K&A . ; ) $ # ; / .(2F ; *. / ! AE /F+(GHF , 2 /A 6 2L F+F. ; , $% 1 ;
, , 2 # # , ; 1 , ; ' $% , ; , 1 D# ; / .(6 +F. +A. (..HF 5
; . 7 5 ; , ' $%
; ; I; ; ; + $% - I
FE 6(.M .
* +
#
, =!
/ .(2F =! *. / = AE /F+(GHF =, , 2 /A 6 2L F+F. =I
, $% $% 1 =I
, , + $% ) 2 #
= , ; # ==
, # =5
, I $% 1 = ; / .(6 +F. +A. (..HF 5!
; $N 1 5!
; , 1 3' 5
; ; A ' $% 5;
; A ' $% ? 55
; I ; + $% - 5
& # P 6 $% ' 1 ,
& # ,P + $% '
/ E & 1 .
B *C ,
& # ;P / $% Q ' "
" / E)& 1
' . B *C ;,
& # P '- " '
1 / E)& 1 ;;
)
& # P 2 ' $% / E)& 1
' . B *C
& # ,P / ' $% ' # 1 1
/ E)& 1
I
& # ;P 2 ' ' - / E)& 1
$%
& # P . 7 B " C ' 1
,!!! B C ,!!I B C I!
& # IP < ' Q Q
/ E)& 1
,!!! B C ,!!IB C I
& # P < ' Q
'-" D# ,!!!B C
,!!IB C II
& # =P / $% '- " '
1 B ,C / E)& 1
& # 5P + $% ' "
Q ' / E)& 1
,!!! B C ,!!I B*C ;
& # P + $% '
-? '- "
' Q / E)& 1
,!!!
& # !P + $% '
-? '- " ' Q
/ E)& 1 ,!!I I
*
& # P 2 ' $% / E)& 1
' . B *C =I
& # ,P < ' " "
/ E)& 1 ' .
B *C ,!! ,!!I 5
& # ;P + $% ' / E)
& 1 ' .
B *C " 5,
& # P / ' $% ' # 1 1
/ E)& 1 ' .
B *C "
5,
& # IP < - ' B C RS B C S,
B C 2#S; B C "
1 5 & # P < - ' .* B C B C < B C 2F B C
"
1 !
& # =P + $% ' "
7 "
) B C 1 B C ' B C !!
& # 5P + $% ' "
7 "
P / $% '- " ) D
/ E & 1 '
- ,
,P / ? - '- B/ C #7 B/4C
B/&C> T . U V " BWXC>
? B@C T . ' B C '
1 ;;
)
P / $% '- " ) D
/ E & 1 '
- I
,P < ' Q D# '
'- / E)& 1
,!!! B "
' Q C I5
;P < ' Q D# '
'- / E)& 1
,!!I B "
' Q C I
*
P + $% ' "
B 13 - C ' "
B - ' C / E)&
1 5;
,P " / E)
& 1 5
;P " 1
/ E)& 1 5I
P " ' / E)
IP < - ' ?
1 55
P < ' Q D# / E)
& 1 "
B "
' Q C ,
=P < ' Q D# / E)
& 1 "
1 B "
' Q C ;
5P < ' Q D# / E)
& 1 "
' B "
' Q C
P < Q B 13 - C '
" "
1 ' / E)
' 3 7 D# " # $%
# ' #
3 "1
# Y #
Z $%
' ' ' # F "
' ' $% '
# ,!!! ,!!I " $%
3 ' ' 1 " #
1 # ' Z & 1
' . B *C ) - '
,!! , ) D ,
' " ) D ? !
7 ' [ ≥ ! " "
'- ' !)
,! ' Z 1 ?
3 & 7
" # ' ' $% P 1 '
& ! " ' ,I '- =
' " =! " ' ,=
'- I # " 5
7 '- ' " ' ?
" " " Q ?\7
$% 1 " ' ? ,!!!
,!!I ' ! =; ] ) ' I= ] )
-= 5 ^ + " ,!!!
,!!I , ^ ' ?
-; I 5 2\ # '
' F " '
==I + '- = ^ '
D ;; ^ " 9 "
I ^ " P ' '
2 * # '- ? "
" P 5 7 B, I ^C
5 2 B , ^C 5 2\ #
B5 I^C '- ? % "
" P 5 2 5 2\
# 5 7 8 " " " B* # C .
" ? A<A
' 2 '
5 7 Z
'-) 7 " F ?
" 1
" ? ' '
" D A<A
A ' 3 _ ` ' a #a " #
# ` ` _ # #
' a 3 # " # #
# a ' a # #
#
_ #
a' " ' _ ` _ a
' ' ' a #a a
,!!! ,!!I " 3 )" # '
" # " # #
1 ' a . B *C # '
' _ ,!! , # a
, ) " # _ # #
# [ ≥ ! " `
_ . ' B!),!
' C _ ' a
3 ' ' V _
# a ' P
' V ' # ! " # # ,I '
= # =! " # # ,=
' I # 5 7
_ ' " ' _ ' _
# " a ? a
_ ,!!! ,!!I ' #
! =; ] ) I= ] ) = 5 ^ F
" ' # a ,!!! ,!!I , ^
_ ; I
5 2\ # ' # '
a " _ ==I F
' =^ ;;^
" & '' I ^ "
' #P ' ' 2
* # ' # "
' # _ P 5 7 B, I^C 5
2 B , ^C 5 2\ # B5 I^C
' _ _ "
_ P 5 2 5 2\ #
5 7 8 " " " B* # C .
" A<A _
' 2 '
5 7 _ ' _ # a #
"
" " #
' a ' "
_ # A<A
1 '2( 3, 4 5
6 " ! ! 7 6"
7 1!
&8 1 " -$
8 $
$ 6
! ! '9: ;<*
.. ,
= >
; #
$ - " -$ $
68 6 6 1
? $ ; )
@ " @ A @ " # 6
& 2 ) ' 9; 9* 3,
> > 6 *
& > " 8
&8 > " B
> > ! 3 6
+ ? '#; ?= ..+,
>
$
6 ! 6 ! 6
= -$
" &8 C 6
7
'2* 9*A .. ,
> - >
6 + > 9
&
& 6" ! $
& '2* 9*A .. D ? 2@*EE= ...,
8 F >
-5 1 - " A
! " ! &8
1! ' , 1! ' - ,
& -$ 'G*; 9*A .. ,
? H > 5
! & ! & 1!
1 6 $ & ! !
>
? 2 * 6 G ?
'9;:2= 9 ,
? - > !"
$ & B ! "
! ! 1
> < 5 !
& 7
! 7 6" B $ " F
" & 6 B$
$ 'A=:<= ,
' ,
$
&
> " 6
! 6 & 6 4
&
" '22 ,
! " #
& ! "
$ ! B B
5 &
4 & ! "
I
! '? ;J EK= ../,
A $ ! >
! 8 &8
$ &8
I & = 6
&
B
$ $ "
&8 1 & ! "
! ! 8
> > !
&8 B
9 ; ' .. , 7
6 & !
6 6
5 $ 5
> 8 & $!
!
8 ! "
-E ' . , ! "
! 1 5
> " !
&8
H >
@ ; ' .. , H >
5 B !
C & $! !
6 B
!
9 6 & ! "
# 7 H @ ' ,
! " $ 5 5 ! $!
" ; " L$
! -G 6 ' M, & $
& 1 " ? '@*, &
@ HB ' ..3, &
! " ! 7 $ 9
' . , > !
" A 2 # J '@*,
* ! ! ! 6 1
? ! ! 5 ! "
I 5 > ' 9; 9* 3, !
" $ 6
&
H ! " > !
* $ N ';@@ ,
! & 1 6$
HB ' , ! 5 ! " I
* 6 5 ? 6 O # ' 3, 5
* & * 1 A 1 ; ) !
3 ! " M+ > L$ 3
3 > 3 7 ! "
A O A ' , >
7 ! " ? * 6
! ! " H >
! > * J
5 1 - ! &8
J$ ! &
B &8
B ! & $ '(A*; K )*
,
@ 2 ' . ., !
&8 >
& ; ! - &
! I 6 1 & &8
1 - <
J &8 G ! ? 7
G
& & > !"
$
& > $ " '0:KE2
.//D G*;;*(; . ,
# - B
! &8 - 1 I >
! " F 5 ! $ $
')=EG ;( O ? A*; .//D 9;:2= 9 D ,
? 7 &
-$ !
! F ! &8 ! !
& ')=2*A . ,
= ! " ! 1
-$ > ! $ 6
$ ' ; PL= D @*;*(; D ? 2 ?K= D
E?=G=; 9=-G(EK= MD 9; 9* 3,
$ % &
= $
! F ! ! ! &
6" @ 5 ! ! &8 >
! !" ! 7
5 -$ #
&8 " 5 $ - 5
&
I '2*E= G(EK= O A=:Q , < 5
5 7
B $ &8 " *
> - !" -$
M '2*E= G(EK= O A=:Q ,
? - > >
6 - "
+. &8
# -$ $
./ + $
1 ' #RA #*;
.. D # ;#=A D 2( 3, = -$ >
> H J
'(#)* +, * - *
* 2 ) 8 # 6
A @ @ " ; ) ? $
@ " $ 1
G & 6 ' =):*(; ..+,
-$ > $ >
" 6 $
& - $
- *
S7
5 ! - 7
-$ 8 B >
" ! 'L ?=2( * .. D ;*#=:T A ...,
9 !
! " K
- $ $ '?K J*A O 0( L=
. /, * B F F 6 F
F ! !
-$
L ' .. , *2#; @ ' ..., C E ' ,
E 1 ' . , ' + / , E ' M M , @
' 3 , U 5 7 '. M , ; " '+ +
, ? 'M , G H ' , J ' M ,
?6 ' 3 , ) ' ,
. 1
6 " 1 !
B !
*
'@ :9VG =V(# 2 V:G@#V)=J @ ; W# ..+,
& -$ *
@ ? E
'@ :9VG =V(# 2 V:G@#V)=J @ ; W# ..+D A:9*2 +,
' ! ( )* #
= # 8 6 B &
2 1 "
& ' , 3 3
$ ' , ! & 7
! $
5 M 1
! $ &
& '22 ,
A E ' 3, +/
&
+ .3 + ?
H
? * @ " &
5 3 6
$ * * "
$ - ; @
@ N ';@@ , 'A:9*2 +,
;@@ * @ " ! 5 3 6
$
I '(# 2 ,
& ! ! &
& -$ $ &
* - / B$
6 $
H ! F &
'0((EE 3,
! & & $
>
! * >
F $ $ -$ ?
$ !
@ ! &
B & $
> &
&8 ' E#:U:*;U:* O 9; 9* ,
> I 1
N 5 * !
> -! ! 1
B 5 5 &
*
! "! - H B $ >
- 6
'0((EE 3,
@ > N - &
6 1 & 1
" & N
>
6 $ ' 9;
9*-E(2 . ,
*
# $ 2 I ?
? > " B >
!"
6
H
" ! !
6 '? A<*EE*<<( +,
=
! $ H
5 & ! 1 5 B "5
+ % , %
#XA #*; 9 " ! $ # ( C EJ ;*Q
J J K D G= <*A E * G D G= <*A 2 @ G = "
! $ # J & C
A#?AD :GJ 9@A ..
-9; 9*-E(2 9 <6 !( ! #!
- . + +.- 3M .
E#:U:*;U:* : @ O 9; 9* E K ? ? 6 I
& $ *
@ # & . M
/M-3
E?=G=; 9=-G(EK= G ) D A 2@ (= * J A # O ;=9 E 2 L D
G " ! & !1
6 1 ? @ & . /
/-M M M
2=;(2 ( E D A 2@ (= * J A # D ; PL= * E G
& - 1 $ A 1 ;
# & . . M 3- M 3
9; 9* E D @*;*(; ( 2 D E*(<* : < D # ;#=A 2 ; J $
! ! !
6 1 " A L ? *
@ " !( ! ! M 3M- 3
; PL= * E D A 2@ (= * J A # D ;=9 E 2 L ? &
! " ! 1 7 $ @
!( ! #! . 33 + 3.3- / ..3
; PL= E J ? !( ! " / # ! ! (
0 1 # 0 2 1 # 3! # #14 Y A
< 5 6 Y @ " @ C * * ! M/
# ;#=A 9 J 5 # ! #! 66$ %! 7, #
8 #! ! 9 A C A*( 'A>
3 ,
#; ?= A 2 8 5 ! ! : ! ! !; A
@ C 2 ..+ 33
? 2 ?K= ; ) J 1# " " 4" # # %! #/7
)* ! / # %! #/7 M ! < Y
? 2 ?K= ; ) J O # @<(A< ) 2 2 $ $!
5 F &
& C , * & * 1 A
1-*A*?V; V# ( C A(2@ZA(= #; A(E*(;= 9* A* A=;( 2* <=
;*2=<= 3 ) I ) I 3
-? 2@*EE= G D ) ;()E(= 2 D A(EJ L D E* E 2 /
" ! # ! * #! ! # " (# 2 ... '# > ,
? ;J EK= L = @ 9 I !
& B ! $ 8 1 #! !;
" ! 1 ! 4(! C <1 B ! $
? -@; *2#; @ V? @G 9 M+ 3M ../
? A<*EE*<<( ? K 2 D A(EJ L 2 ? D < # ;*EE( 2 D A <=A
2 2 + U [ :
( C L 2 ? A 2 < 2 < G O E J E
' , # (! # #! # 8 4 ! ! )<! 5 4 5 ! ( )* . - 2 > 2 # "
?K J*A E K ) D 0( L= < ; & V
$ 1 -$ !( !
#! 9 # % M 3/- . /
9;:2= 9 2 0((EE E K @ E(2 @ ? G =E(J*(; 2 ?
=E(J*(; J ; E#:U:*;U:* A ) A?(2* <= ? * A O
? J E? <* L * > $
( C A $ & ! & &8
$ & 5 & $ &
!" ? =
*2#; @ V?@ <A :G@* ? ( #
@
9;:2= 9 2 D E(2 @ ? G ED A=:Q A 2 A
> ! A 2 # J Y @*
! #! 0! >1 ! . + +/-3. .
9: ;<* ; <
6 -$ C & #! #! 1#
% . + -3M ..
*2@;*A #; A(E*(; 9* @*AU:(A );=@*?:\;( -*2#; @
% ! & ! #! " )* #! ; L C
*2#; @ V? @A ... + '*2#; @ -A 9 3,
G*; 9*A # A -$ !( #
G*;;*(; ; E ? 4 ! 1 1 # (! ! )* # )* ! #! ?5! ! )* #! )17 . 1& # &4 5
!; " ! . . ! 9 & 2 Y :
G J & J &
)=EG ;( E O ? A*; ; E @ ! ! ! / # ! *
#! ! 5 !?5! ! )* " ! # K 5 C
@ :9VG =V(#9GV#; +3 .// 'A> <> ,
)=2*A 2 G = 6 C ? $ A! !
M . +
(# 2 # #! #! ! ( )* 8 ! ! ( 5 1 # 5 2
1 9 " C ] 6 CVV^^^ V_ C
B
( A<(<:<= #; A(E*(;= 9* )*=); G( * *A< <WA<(? -(#)* B 5
#! # 8 5 ! 5 ? )* 9 ) 7
+
(A*; K )* ( " " ! 0 4 !
5 #! ! 15! )* #! 4 ! #! # # 8 1 ( )*
/3! '9 & 2 , : G @ $
L ?=2( * @ 0 < % & 8 ! ! 1
( C EJ ;*Q J K D G= <*A E * G D G= <*A 2 @ G =
" ! $ #
J & C A#?AD :GJ 9@A .. .3- MM
0((EE E K @ 8 5 2 & ! ! ) #
9 " C
]6 CVV^^^ V V 6 [ ` _ C
M 3
0:KE2 * J & ) ! # ; !(
# .//
3-E* E ( ; D A(EJ L 2 ? D < # ;*EE( 2 D E ?K*; L; < * 2
& ?
# B! # (! # #!. 3
E=@*A @ ; ? ! ( #! !; #! ! 4 1 5 ! 7
4 # & ! 9 " C
<^^^ V V V / 6 _ C
. 5
2 ;<( A G ; G ! # C 6 1
2*E= G(EK= L G O A=:Q E J = B &
A -$ C ! !( :
/ M
2* 9*A # J = A -$ ( C ?= );*AA= ?(= E
A=#;* *AAa ?( A <(J A .. A @ A @
.. M.+-M..
2( (A<4;(= 9 ( <*); Tb= ?(= E-2( ( #! )* #
%! 74 # & ! # " 2( VA $
; # " 3 MM
2( (A<4;(= 9= 2*(= 2#(* <*-22 ( )* ! )<!
5 4 5 ! ( )* # & # (! # #! #
:G@*VG & 9 V? &
( # VG & # V*2#; @ - -$ C
22 VA#G +
2( (A<4;(= 9= 2*(= 2#(* <*-22 , ! 5 ! # 8
# #! #! ! ( )* 9 " C
]6 CVV^^^ V ! [ `V V !V V
V 6 O ` @ _ C B
=):*(; 2 !# ! ! # ! * ! 74 # #
#! ! # ?= G*;a ?( ?(= E * A*2( \;(= E <( =-2*;(? = 9 9*A*;<(G(? Tb= G 5 C ..+ /
@*;*(; ( 2 D 9; 9* E D ?=A< L ; 2 D 9( A L 2
; & !
" & @ & 4 . 3 M
+ M-+
@*;*(; ( 2 !( ! " # ! &/ ! ! 4 !
# ! 1 1 " / #! ! ! #! &
# "! ! ! (! #! 5 / 9 & '2
@ & J , - : G @ "
@ :9VG =V(# 2 V:G@#V)=J @ ; W# # !
" ! # 0 & L @ C @ :9VG =V(# 2 V#; V /V /
..+ +
;*#=:T A @ $ I -$
( ?= G*;a ?( ( <*; ?(= E 9* ? @< Tb= 9*
\): 9* ?K:J . ... @ ...
;=9 E 2 L # (! ! )* &1 ( 7 &/ ! !
>1 4 ! #! ! 0! &1 .. . ! '< ,
A < L A O A=:<= L A 9
! 1 * & * 1 A 1-; !(
#! ! 9 # ! . MM- +
A=:<= @ ? 1 1 )* ! #! 5 )* #! ! 5 :! !
# &1 )* #! !#4" ! 4 ! #!
0 & . 3 ! < '9 , Y
: G @ "
A:9*2 1 3 )* # # / " ! # # # 0 &
L @ C A:9*2 +
<();* ? # 0! >1 ! !?5! ! )* " ! 5 3 !
" # $ % & "
' & % () * +
,-- , % + . /,
& + .
/ - 0 12) ≥
$ /3-4 (/,53
4-3 * ,3 ,6 /7
89 2 2
: /337 ;
<3 = .
66 = > 74 =
? @ 1 8
2 :
? & (,/ 7 =*
8 (/, 4 =* 89 2 (A 7=* 2
? 8 89 2
& ( * '
&
C
C %
& ! D E D C%
& "
C ' & % () *
D ,-- , C /,
2 + D /
0 12) ≥ /- D
; /3-4 (/,53 4-3 *
,3 ,6 /7
89 2 C &
D /337 <3=
66= $
74= ? @ 1
8 &
D ? & (,/ 7=*
8 (/, 4=* 89 2
(A 7=* & D D
D ? 8 89
2 & ( * '
G H / <-- --- E ,
/A= I
(J K$2LM N 12'@L /533* @ "
+
+ " ; %
+" 53- --- F , // 4 =
. (;O2L&@ /55,P 8MG ,--7*
2 %
H . 2
. +"
" "
(;O2L&@ /55,*
' Q CD (/553*
+"
2 @ "
<-= G (2G;L2;@ ,--7*
.
;
+" H
# #
. : 4-- 2
+" %
+"
# (;O2L&@ /55,*
2 +K (/5A/*
#
. " "
% ;
+" (&2R2L@'
/537P L ;2K /55,P '2G& ' /55,P 2L2ST /557*
@
"
!
! "
(' OU2 ,--6* 2
" ! !
% V
B
(12LR2KQ /553*
2 "
!
" L))G+$ % & "
#
!
"#"$%&#'(")*+ ," -#&" ,& &.%/,+
" # $ %
& " -3W --X ' 63W ,6X Y
8 . 2 " K % ' & %
() * /A E ) ($ /* & +
L ) ) G L))G & "
) (GW//-Z5A+G* M +) 2 "
6,7 "
. 2 "
H
0 K % "
' & % @ )
" + H '
) (' ) *
,7-F[ .1 +"
2
<-- 2
+ + ! ( L2'MK /53A*
; B (T
,--4 T ,--7* %
343 3 #
+ . . % 2
+ ! !
2 " % . ;
' ! H H +
%
H ('O;@82 ,--4*
. "
) +1
# #
) K 2 G K .
) (@8 L2)2 /555*
2+.%#"3&2 & $+4&%" ,& ,",+. ," $+25+.')*+ 64+#7.%'$"
)
% 4A " H (,-- ,*
" % L
(/55,* "
$ H
+
7--
0 ; (3* (4A*
L))G $ % & " ' & % () *
$ % /,
B ( N $ /55,* 1
.
/
-/- - 2
. 2
B %
Transectos que cortam a reserva.
!
H
Q " \K ] ^ O $
) 1 T ) () *
2 %
1 (/5A/* +
"
$ ? % "H
(L&* Y E (/537* '
(QX* ) (T* ' (1*
D N U (/5A4* ) % +
. ?
• _ % "H
( )
N
T
RT
ln
=
?
` G: "H P
` : P
` K
• _ ' (QX*
( )
( )
N
n n N
N
H
S
i
i i
−
= =1
ln ln
'
?
` : P
` : + P
` : P
` ( *
(
)
(
1
)
1
1−
−
=
=N
N
n
n
l
S
i
i i
?
` B
` B ' P
` : + P
` : P
` :
• _ ' (1*
max
'
'
H
H
J
=
?
` @ ) P
` ` "H P
` : ` %
! 8 G R , -4 % %
8
2 + . / 3-4
,3 ,6 /7 (& /* ;
3< = ,4 = :
+/ /337 1 " ,3
<3 = . 66 =
0 L + . L))G
$ % & "
"274'"9 .5:$'& +2& ;/43"# <-='%+
2
" # 2 2 2 .
2 C
8 ) 2 .
T 2
$ (8 H ;1 * ' ) a 2 .
% $ (' Q T
N 1 * 2 L C @ 2 .
(8 * T J M 2 .
1
& 1 2 .
8 @H & T " 2 .
( * ' 8 . 2 .
(1 * M D N
C ' T 2
1
(K * K $ ! 2
1
$ ' Q 2 2 .
1
8 8 2
@ C HC
& $ @ ' % L 2
@
(8 2 * ) H F
Q $ 2 .
" $ ' # 89 2 8 2 .
89 2 8 2
& 8 ) 2 .
89 2 ) 2
& / 0 1
$
(2 * 2 1 ' 1 2 .
8
2 2 .
% ( * ; E T 2 .
" (Y * ) T 2 .
" 8 H T 2 .
) C
8 1 : 2
L
( ' ) ' 8 T % 2 .
@
2 :! (,---* " +
1
2 ?
T"
" $ ' #
% $ ( ' ) '
$
G . /A
// 8 @
1 :
G
@
%
2 : ?
, 2
@ %
' . B
: (2G;L2;@
KM82 /5A/P 12821Q ,--/*
K N L (,--,*
' 1 )
@ 1 8
63 7 =
1 4A
" $ ' #
8 H :
2
$
% $
( ' ) ' T"
' 2 (,--7* " #
"
" % @
"
> 74 =
($ 6* 2 :
? (,/ 7 =*
(/, 4 =* (A 7=* ($ 4* 2
:
('28)2M /55<P 8@OGM@L N 12LR2KQ ,---P )@L@ML2 ,---P
2L2ST $MKQ ,--,*
0 50 100 150 200 250 300 350 400 450 500 Caes alpin iacea e Comb reta ceae Mim osac eae Euph orbia ceae Apoc ynac eae Burs erac eae Faba ceae Eryth roxy lacea e Capp arac eae Bign oniac eae Anac ardia ceae Polyg onac eae Bom baca ceae Coch losp erm acea Rham nace ae Famílias N ú m er o d e in d iv íd u o s
80 L B :
L))G+$ % & "
0 100 200 300 400 Caesalpinia
pyramidalis Tul. Combretum leprosum Mart. Croton sonderianus
Müll. Arg. Aspidosperma pyrifolium Mart.
Anadenanthera colubrina
E
sp
éc
ie
s
Núm. indivíduos
>0 @ :
" L))G+$ % & "
_ ' (QX*P @ (T* _
' (1* , 63 +/P - 3/ - A<
(& ,* @ .H #
%
/ 5/ 6 -5 Z (2K1 $ L2; $MKQ ,--6P 2L2ST
/557P $ G'@12 /55/P KbL2 /5A,P )@L@ML2 ,--, ,--6P
L ;2K /55,P L ;2K /55A*
0 L % (L@* (LJ*
(L$*P _ ' (QX*P @ (T* _
' (1* "
<? @
6 <, 6 -5 , -/ , 63 - 3/ - A<
@ " ' . L J G
1 K (/555*
% ' T 1 () * +
L))G+$ % & " V
QX "
%
.H ) (,---*
"
6- ' H
% #
# #
2 "
H 2 :! (/557* L
(/55,* 6 <3 4 37
<-> A
* 2 + +
.
% P
* ' (QX*P @ (T* _ '
(1* % H
H "
. P
* 2 " " "
H :
% P
* 1 4A
" $ ' #
" P
* 2 : ?
B C
2K1 $ L2; +$MKQ $ J P '28)2M @ R ' P L ;2K 8 T G P
$ .
. 1 ) $%" =+% #". /3
, ,A3+6-6 ,--6
2G;L2;@+KM82 ; & &;'.%" #".'4&'#" ,&
+%DE'$" 4 /45+/76 /5A/
2G;L2;@ K 2 P )@L@ML2 M 8 P K@M&@ O & P 2L '2 8 L R 2 "
. ' T 1 @
) &;'.%" &#E& // 6 ,76+,<, ,--7
2L2ST $MKQ T 2 P 12LR2KQ $ 1 P J2L1M2 L @
% "
&;'.%" #".'4&'#" ,& F++%&$E'" 6/ / //+/5 ,--,
2L2ST @ K P '28)2M @ R ' P L ;2K 8 T G 1
. " 1 )
&;'.%" #".'4&'#" ,& '+4+3'" 77 4 757+<-3 /557
2L2ST K R 1 &;"E%"2&E%+ '%+..+$'+4G3'$+ ," &.&#;"
"#%'$/4"# ,+ "%#'2HE'+ "%/#"4 ," "(&E," "2"E,/- 0 '
& % 0 ) ) ,--- 63
L2G1 ' 8 0 5"'."3&2 & + 1+2&2 .&#%"E&I+ '
) ? 8 /554 77
LM;' G ; P $ L82G K /55, 1& 1&#="#'/2 1"E,=++J L C
J K
L Y@L T @ P U2L T Q '&4, "E, 4"=+#"%+#K 2&%1+,. 6+# 3&E&#"4
&$+4+3K ; ? Y 8 1 D /5A4 ,,<
12821Q L J R .%/,+ 6'%+6'.'+3#-6'$+ ," $""%'E3" ,+ &#',G
.%")*+ &$+4G3'$" ,+ &#',G ,--/ /6- & 0
O ' )
128)@KK $ P J2LMJKM 8 2 P 'MKR2 T 2 P K@2K 2 8
'"3EG.%'$+ 64+#&.%"4 ," #&3'*+ E+#,&.%& M 282 /555
( *
1L G>OM'& 2 " 'E%&3#"%&, .K.%&2 +6 $4"..'6'$"%'+E +6 64+L&#'E3 54"E%. G D b E 1 O C ) /5A/ /,<,
;O2L&@ L &
+" ? ",&#E+. ,& .%/,+.
+$'"'. 5 / 4/+76 /55,
$@LG2G;@' 2 ' +" G &;'.%" ,+
E.%'%/%+ 4+#&.%"4 4 //5+,4 /55,
$@LG2G;@' 2 ? M ? 2L2OT
@ K ( * '+,';&#.',",& +E.&#;")*+ & .+ /.%&E%-;&4 ," 4+#"
,+ #".'4 L )@? @ ,,+ ,< ,--,
$ G'@12 8 L /55/ E-4'.& ," ;&3&%")*+ "#=/.%';+"#=G#&" ," $""%'E3" 1'5&#M&#G6'4" ,+ E+#+&.%& ,+ &.%",+ ,& '5& &
; O @ 1 1
J K$2LM K P 12'@L L K F+E&"2&E%+ &$+4G3'$+ ," #&3'*+ +#,&.%&
5"#" &M5&#'2&E%")*+ 64+#&.%"4 Q % ?
)GO;Z$2 ZM ;$Z L2 47 /533 /<< (' & /-*
Q@bY ; R Q &E%#&. +6 54"E% ,';&#.'%K YY$ZMO1G K /553
MG'&M&O& L2'MK@ML ;@ J@ JL2$M2 @ @'&2&_'&M12 0 M J@
"5" ,& '+2". ,+ #".'40 5#'2&'#" "5#+M'2")*+ ;
J ,--4
K@8 ' T L P L ;2K 8 T G $
$%" =+% #-. /< / ,6+4, ,--,
KbL2 2 K L & /5A, 6&'%+ ,+ #&4&;+ E" ;&3&%")*+ ,& ,/". -#&". ,+ 2/E'$75'+ ,+ #&I+ ," ",#& ,& &/. ;
8 O $ L ) L
8@OGM@L M 8 T P 12LR2KQ 2 T @ #&.$'2&E%+ ," $""%'E3"
./=2&%'," " ,'6&#&E%&. %'5+. ,& $+#%&. E" #&3'*+ ,+ &#',G ,+ '+ #"E,& ,+ +#%& G ? ) ! 882Z$2 ZO&$Z L2Z-43 ,A ,---
8MGM'&VLM ;2 MG&@JL2cd G21M G2K ,--7 +;" ,&4'2'%")*+ ,+
&2' -#',+ =#".'4&'#+ 8MGZ' "
)@L@ML2 M 8 &;"E%"2&E%+ 64+#7.%'$+ ,+ &.%#"%+ "#=G#&+ & "E-4'.& ," &.%#/%/#" 6'%+..+$'+4G3'$" ,& &$+..'.%&2" ,& $""%'E3" .+= ,'6&#&E%&. E7;&'. ,& "E%#+5'.2+ ,--- 3- ; (8
) R * + O $ )
LMUUMGM 1 & #"%",+ ,& 6'%+3&+3#"6'" ,+ #".'4 ,e f
1 @ K L T /553 6,-
L ;2K 8 T G '%+..+$'+4+3'" ," ;&3&%")*+ "#=/.%';+ "#=G#&" &2
N/"%#+ -#&". ,& $""%'E3" &2 &#E"2=/$+ /55, /5A (& *
O @ 1
L ;2K 8 T G P 2G;L2;@ F R ' 2 P '2K@' 8 $ P J 8@' 2 ) '
$ :
) &;'.%" #".'4&'#" ,& '+4+3'" 7A 6 7/3+7,< /55A
'28)2M @ R ' $ M ? '28)2M @ R ' P 82b ' T P
2L '2 8 L R (@ * &.N/'." =+%DE'$" E+#,&.%'E"0
L ? ' B Z' L
) /55< ,-6+,6-
'2G& ' 8 $ 2 R P LM @ML 8 L P '28)2M @ R '
' &.N/'."
3#+5&$/-#'" #".'4&'#" ,3 6-7+6/4 /55,
&2R2L@' ' P )2MR2 $ 2 $ P &2R2L@' @ T ' E;&E%-#'+ 64+#&.%"4
E" "#"7=" & '+ #"E,& ,+ +#%& @
R ) L ? 'O;@G@ /537 6/
. . # .
/ 0 ( 1 2 3
-++ # ! / -4 #
-4 + # 4 /
-! #
/ # 5
6 57 "
5 !
- 42 8 .- -- !4 8 .- / 4 , 9 0
# ! : 9
5 / 2 -!
#
/ ; "
"
; = > ? ?
> > =
? @
? ? ? !
? $ % & '()* = +, -
? . >
= 0 ( 1 2 @ -++
! -4 -4 +
4 -!
= =
= ?
5 ? %
= ! - 42 8 .- -- !4 8
.-4 , 9 A ?
! : 9
= 2 -! B =
%
A / D
"
57 A
." E
# &
/
5
6
!! 9 - !+, F4 > "
';)GH +* I" 5 5
J / '0KLKH -:, M
G;KN;H%%H +* /
." & .
J
# 5 &
( . 5
#
"
." #
" ."
5 &
" H
# #
J 5
& D
6 O " '(HBH;B
D #
" /
#
E
" 5 "
/ "
" J
. P
H
" #
B '-:: * 5
" &
/ D "
6 5
5
D #
A
"
! "# $%& ' ( ! '! !) *'&
" " B ( ( P
E 3 & % " O Q
" N # $ % & '()* -,
> ( '3 -* " . .
) $ ( ! 2-
4R S $ 24R 2S @ '3 *
"+* , N & " #
$ % & H ( #
0 " CT
" & . )- ."
')B $;N -:4,*
& . .
5 " "
/ ' B UVA * ) .
;)GH '-:: * # " /
$ /
"+* , B " " B((E.
3 & % "
& "
" & 0
$ / #
. & D
5 #
'$K0HQ +*
! &'&.&+" '! /&) +!/ ! &.! '! ' '&)
E +,
5 " ↔
D - # "
6 -! '3 2*
5 5 E
# ' $*
' (*
# 5 -
# - #
"+* 0, Q / B((E.3 & % "
# /
" "
D ! # % "
5 #
( 5
6
0 1(.")! '&) ' '&)
0 &/2&)"$%& 3.& 4) "
6 &
W " XN Y Z K 3
( # V ( '()* 3
5 '-:,-* (
& &
'-:2,*
0 1(.")! 3" &))& "&.5+"
3 &
! "
& /
& & "
6 6 57
/ 6 5
/ O
& / D J " "
'N Q(BH W% -:F+M 3;EAN -:4-M QKHNNHB.0KQ)A;$ [
HNNHE)HBG -:4+*
A " 6 "
& . D <
• 0 B
100
1
x DA DAi DRij
i=
=
\ 0 '9*M
• 0 6 B
100 x AB ABi
DoR=
\ 0 6 '9*M
\ ] # /
<
4
2 π
x D
AB= M ' \ 6 *
• 3 57 B
100 x FA FA
FR=
\ 3 57 '9*M
\ 3 5
• I ; 6
\ 0 '9*M
\ 0 6 '9*M
3 57 '9*
( 6 #
" # / 5
6 E 6
2 #
< /
22 2 9 # 'BA0 N -:: M
$AK$ [ NH;%H -::2*
0
0 *3" "61 " /&) .
/ ' +* . 5 J
/ !
O 5 "
& &
, ,! "
/ "
"+* 7, $ ' * "
' * ! ' * 0 5 10 15 20 25 30
0 2000 4000 6000 8000 10000
Área amostrada (m2)
N o es p éc ie s ac u m u la d o 0 5 10 15 20 25 30
0 2000 4000 6000 8000 10000
Área amostrada (m2)
N
o e
' -* 5 & B
G E " !2
H & $ [
$ ' F* / / +
5 # .
& 5 "
F #
0 &/2&)"$%& 3.& 4) "
3 -++ # ! /
-4 # -4 + #
4 / -! # '% -*
, B # / . B((E
3 & % " /
#
/4." 8 )29 "! !: 1 /!1 & ;(<" & === ==>
Y Y
?
Q Y Y
)
V 5 Y Y
! ) [ W > Y .
! " 'Q 0 * $ Y Y
)
# " '$ W V [
* B ? Y Y
)
$ $ 'Q * V ) G Y Y
'N * N Y Y
$ " $ W Y Y
Q Y Y
H ? ?
% $ " A H $ & Y Y
H
$ $ 'Q * ( C
W
Y Y
$ " & Q7 . Y
% - _
% $ $ $ Q . Y
' $ Q7 Y Y
$ W Y Y
% Y Y
Q H % Y Y
$ $ $ ') * $ Y Y
( $ ' * ; = [ ) ? . Y
3
' * $ Y Y
Q
$ Y Y
# ') * 0 > Y Y
'@ * ( Y Y
$ $ Q ) . Y
E?
) Y .
A
* N Y .
( ?
! $ Q . Y
B
+ & $ , & Q Y Y
0 Y . .
E
-, / # -2 # 5
Q / A #
'-+* # /
X Z E -, /
# -- # 5 Q
H 5
J / 5 #
0 / ) *
- 2 # # E?
A &
! ; #
5
# % $ " #
H ? ? % $ $ $ # H
#
." #
D # 'BA0 N -:: *
E 6
# #
Q H
'3 !*
6
J # /
# 5
B /
D 'LKH;BA` -
"+* >, I 6 # 6
B((E.3 & % " ' * !' *
A 6 "
<
<
$ '3 F*
6 - - ' !*
"
5
6
E / #
# /
# # H
b 0 5 10 15 20 25 30 Caes alpin iacea e Mim osac eae Euph orbia ceae Comb retace
/ O # 5 & 5
# E / 5
5 & 57
/ 6 # H
" ' N)KLKHBLKH -:, M BA0 N -:: M B UVA
- -::!M 3HBB ` -::,*
"+* ?, I 6 /
' * !' *
0 0 ) * * -& "#&1 .
A J # 5 -!
'% * # b 0 5 10 15 20 25 30 C.py ramida
lis C.so nder ianus C.lep rosu m A.py rifoli um A.co lubrin a M.te nuiflo ra P.sti pulac ea C.lep toph loeos B.ch eilan tha J.moli ssim a Espécies V al o r d e im p o rt ân ci a (% ) 0 5 10 15 20 25 30
C. p yram
idalis
C. le pros
um
A. p yrifo
lium
A. co lubrin
a
C. so nder
ianu s
C. le ptop
hloeo s
M. t enuif
lora
P. st ipula
cea
B. ch eilan
tha
$
, 9 # O /
244 .- !
9 2 9
-! 9
E ! '% 2* -44! #
/ -, 9
O /
$ #
!, 9 # 0 5
/ !
2, .- - 9
- 9
4 9
$ - - '-::,* ( - - ' -* 5
/ - #
5 5 6 /
# H .
J #
-5 / / /
" "
"
0 ! 4 / !
! 9
, 9 J / -+ 9 !
5
"
H O " /
! '% 2* . 5
- 42 8 .- -- !4 8 .- !
#
$
'% 2*
H & B G
E " 5 F
-8 .- -, : 8 .- '3HBBH;B -:,,M QAB;Q - - !M
$ E% E [ $AK%A F*
3 - - ' 2*
.
" F : 8
.-- : 8 .- A 5 .
H & D
, I 6 / B((E.3 & % "
'H 6 *
&/! ' !)29 "! @ & A
B/ C D
% 224 2 !44! 2 + -2 :- 2+ 4+ +
-Q7 2F - - , ! -+ - -2 - 4F F
-Q - 4!- -+ !, -- ,2 4 : -- +
Q - ! - ++ 4 : : +4 : :! , :
$ 4- - ,,+ + :2 4 + - !- , ,
'@ * ( 4 ,! 2 + ,F 4 F: , , F :+
# ') * 0 > :- 2:! F 2 F , 2 ,+ ! F!
$ $ 'Q * V ) G 2! + F +2 ! : 2 :- + :
$ $ $ ') * $ 24 + !4 + 42 - :, 2 :
' $ Q7 + F 4 - F4 2 ,! !: 2
' * $ -2 -!F! : FF - ! - 4
0 -+ +42 :4 FF +F - 2F
$ " $ W ::4 - 2: ,: :4 - ,
% $ " A H $ & - - 44 ,2 ,: - ! :
! 22F 2! - +, 22 4
# " '$ W V [ * B ? ! ! : 2! - -, !- F,
Q H % 2 4- - ,: F +!
'N * N + -42 , ,: -4 ++
! " 'Q 0 * $ + -FF , ,: -F ++
V 5 + F, , ,: 4
+-+ & $ , & Q 2 -2+ - ,: -2
+-* N 2 !+ - ,: ! 2,
! ) [ W > + -+ 2 +
$ W 2 - ! - 2 -
-$ $ ( - -+ 2 - -,
) - 2: 4 2 + -2
77= =E FF7 == == == ==
0, I 6 / B((E.3 & % "
! 'H 6 *
&/! ' !)29 "! @ & A
B/ C D
% 2F! + :2: - + -- F 2F ,F 2 2
Q -- !, ! - 2, , !: ! 2 , 42
Q --4 - ,42 F ,4 4 ,2 : 4: , -F
$ ,! F,F2 + :: , !: F -, F !,
Q7 -+F 2 -+ , !4 F !4 ,: F
-$ $ 'Q * V ) G +4 ! -+ 4F F 2- + 4 + !:
'@ * ( + F! + +F ! ! ! ,F + +F
# ') * 0 > F! 2 : 2 ,- ! ,- , 2 :
$ $ $ ') * $ F2 --+ 2 4 ! !F - : 2 4
' * $ FF - : 2 2 - ,F F
' $ Q7 ! 4F! - +4 4, F: - F+
% $ " A H $ & -F :-- :+ 4 , - 2+
$ $ Q ) -, ! + - F 4 +4 - 4
'N * N - !-! 4 4 +F - -!
4 2:, +- - 44 2F ,!
% $ $ $ Q -2 : 4F - F F 4F
Q H % + + 2 - - 2F !+
# " '$ W V [ * B ? + !!+ 2 4F ! !
( $ ' * ; = [ ) ? ! 42 : - - 4 +F
! " 'Q 0 * $ + -: 2 4F +
$ " $ W + ++- 2 !- + 2,
V 5 + F2 2 4F F 2!
+ & $ , & Q 2 -2 -, 4F - 2!
$ W F -, 2! !- -F 2+
! $ Q ! +: : ! F
$ $ 'Q * ( C W 2 - ! -, ! : -4
$ " & Q7 - - F ! -
--G=7 E => == == == ==
% !
/ 2 4 8 .- + F 8
! 2! 9 24 9 6
D - - -
-- - - $ #- - $ $
'- 44 9 6
E ! - -
-- - - $
-#- - $ $
-,, 9 6 '3 4*
G, B & / "
' * B((E.3 & % "
' * ! ' * 0,0000 0,5000 1,0000 1,5000 2,0000 2,5000 3,0000 3,5000 4,0000
C. py rami
dalis
A. co lubrin
a
C. so nder
ianus A. py
rifoliu m
M. te nuiflo
ra
C. le prosu
m
C. le ptoph
loeos
P. st ipulac
ea
B. ch eilan
tha
J. mo lissim a Espécies Á re a b as al ( m 2 ) 0,0000 0,5000 1,0000 1,5000 2,0000 2,5000 3,0000 3,5000 4,0000
C. p yram
idalis A. p
yrifo lium
A. c olubr
ina
M. t enui
flora C. le
pros um
C. le ptop
hloeo s
C. so nder
ianus P. s
tipul acea
B. c heila
ntha A. c
A "
$ ' !* #
'()* $ B G E
/ 57 5 !
:, 9 :F 9
A / 57
',! 9 !+ 9* ',2 9 4- 9*
'F4 9 F+ 9*
I" 6 /
G & /
/ 5 'BA0 N -:: M $ Q( ;A -::FM
N A3AB 0A.3;NWA - - 2*
5 5 !
57 O /
5 /
0 0 ") "<*"$%& '" /9 " ! !/ . *
0 0 ") "<*"$%& '" /9 "
A 6 " 'F, * 5 ! '2,
* E #
$ 5 / 5
" #
A 6 / # ', *
5 $ ' !* "
$ 5 F F A
J #
# /
V. '3 ,* 5 / #
5
0 -++ # : 9
. 5 / $
/ '2a..F* +2 9
# E J -
# , 9
E !
: 9 # . 5
/ 5 +, 9 '2a..F*
5 , - 9
J
H & )
'BA0 N -:: M (HBH;B - - * 5 /
#
H, 0 #
6 B((E.3 & % "
' * ! ')*
E
# - -+
# 4 9 /
V. '3 :* 5
5 J
# / 'Fa..:*
a 0,0 5,0 10,0 15,0 20,0 25,0 30,0 35,0 40,0 45,0 50,0
3|--6 6|--9 9|--12 12|-1 5 15 |--18 18 |--21 21 |--23 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
In d iv íd u o s (% ) 0,0 5,0 10,0 15,0 20,0 25,0 30,0 35,0 40,0 45,0 50,0
3|--6 6|--9 9|--12 12|-15 15 |--18 18 |--21 21 |--23 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
F, 0 / 5
/ 6 B((E.3 &
% "
E !
$ #
44, # +F 9
J # /
" / V.
'3 - *
Caesalpinia pyramidalis 0 50 100 150 200 250
3|--6 6|--99|--1212|-1 5 15|--1 8 18|--2 1 21|--2 3 24|--2 7 27|--3 0 30|--3 3 33|--3 6 36|--3 9
Classes diamétricas (cm)
N ú m . d e in d iv íd u o s Croton sonderianus 0 50 100 150 200 250
3|--6 6|--9 9|--1212|-15 15 |--18 18 |--21 21 |--23 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
N ú m . d e in d iv íd u o s Combretum leprosum 0 50 100 150 200 250
3|--6 6|--99|--1212|-15 15 |--18 18 |--21 21 |--23 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
N ú m . d e in d iv íd u os Aspidosperma pyrifolium 0 50 100 150 200 250
3|--6 6|--9 9|--1212|-15 15 |--18 18 |--21 21 |--23 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
=, 0 / 5
/ 6 B((E.
3 & % " !
( /
# /
L & - - ' F
0 0 ") "<*"$%& !/ . ))!) '! . *
A /
# '+ * "
'-- * # " / - 5
" #
( "
< '- a...2 !* /
'2 !a...! !* '! !a...-- * E
4 9 J # 4, 9
J / 0 50 100 150 200
3|--6 6|--99|--12
12 |--15 15 |--18 18 |--21 21 |--24 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
N
o d
e in d iv íd u 0 50 100 150 200
3|--6 6|--99|--12 12 |--15 15 |--18 18 |--21 21 |--24 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
N
o d
e in d iv íd u Aspidosperma pyrifolium 0 50 100 150 200 250
3|--6 6|--99|--12 12 |--15 15 |--18 18 |--21 21 |--24 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
N
o d
e in d iv íd u o s Anadenanthera colubrina 0 50 100 150 200 250
3|--6 6|--99|--12 12 |--15 15 |--18 18 |--21 21 |--24 24 |--27 27 |--30 30 |--33 33 |--36 36 |--39
Classes diamétricas (cm)
. -, 9 . -4 9
. -! 9
-9 / ' $ - 9 H / 4
2 9 # , 9
/ 5
E ! , 9
/ 9 # /
.
-9 . - 9 $
$ . 4 9 . F 9
. ! 9 & +2 9 + 9
/
E / FF 9
# / "
/ , 9
. 2 9 . -! 9
# 4 9 ! 9
& , 9 # -: 9 /
E ! / F4 9
# # :F 9 /
. 2 9
. -2 9 . -- 9
! 9 $ $ $ . + 9
& !2 9 # 9 /
/
E 4 !9
# ! 9 /
. 2 9 $ . -!
9 . -2 9 . - 9
. 4 9 & 44 9 # + 9
E ! J #
-2 9 F! 9 / "
/
. F 9 .
-4 9 $ . -- 9
. , 9 # . , 9 & 4 9
# 2 9 /
!
. 5 ! #
7 I
* # 6 5
Q H
M
* A 6 "
E
$ M
*
6
5 6 "
M
* E H O #
5 & # / 5
57 /
6 # H M
* H O " /
. 5 #
/ , 9 6 M
* O 6 : 9 #
5 !M
* H ! #
> J
N)KLKHBLKH $ G M $A BH$ V G G M B UVA 3;NWA V !1)"' '! '! !)29 "!) <*) ": ) !/ :!+! $%& '! "1+
( HQ)B ( . ( %$ '( 5 -F* -:,
N A3AB 0A.3;NWA 3 G M $ Q( ;A H I $ ) M BA0 N Q V E M
3 #
( <& )E -4 ,4.2 2 2
QAB;Q ; N M $ Q( ;A H I $ ) M B UVA H N 3
. " $ BE
) <& )E -: 2 F-!.F 2 !
B UVA H N M $ Q( ;A H I $ ) M BA0 N Q V E
# " (
!:") )".!" '! "&.&+" E !! + !:!.F 4 -::!
B UVA N I !: 1 /!1 & " &))& "&.5+" & ' !)! :
" *. '& "/K1"& * . ' #!1' / 1'*( _ $
% & _ ( # ( 24
)B $;NbQ ) *'&) <()" &) 2 & .!: 1 /!1 & + 4 &. , 2 "'%&
+ 4 &. ' ) ! ) ' 4< ) # < );E GB; 2 -:4, 2
;E $ % . /! " 1 "'. 1' * .")
-:: !42.!,- -:2,
Q WA B G I ) *'& 3" &3")"&+ (3" & ' "1+ '& ! "'5
) $%& ! &.5+" '& ! "'5 - -2 % _
K $ (
$%BA V 3 .& 4) " ! 3" &))& "&.&+" '! */ ! '&
/ +"1 . < )".!" &E L*! ) '* . '! A $*1*1+ E 1 " '&
)) M* & -:,4 2, 0 'Q * K
H .$(
BAELK;$% 1 "1 !+ !' )N) !/ &3 . ))"3" "&1 &3 3.&O! "1+
2. 1 ) E = c > K ? ( -:,- - F
0KLKH V G & '!) ! ! ) . :&* ) P! 53". ) 2 Q <
H$ Q -:, 4F
I ( J ( !:") )".!" '! & Q1" E - - 4.-! -::,
3HBBH;B B N 1(.")! !) * * . ' :!+! $%& ' ) $%&
.& !) . '! P2! "/!1 $%& '! $* E &/& )*<)4'"& <()" & 2
& / 1!R& 3.& !) . -:,, : 0 Q _ K
3 I I
3;GKHBH0A Q !+"%& '&) 1- /*) 1& '&/41"& ' )
"1+ ) Q $/ ) +-- -.2+ -:,2
3;EAN K Q E 6 "
A! & A!1!#E -+
- :.+
-:4-3B E HN;EA Q B M 3HBE E0H$ 3;NWA H ; M BH$HE0H Q M NH;%H W G
. . !:") :& ! 4
- 4:.,F 2
G;KN;H%%H Q 0
; < V Q $;NI Q % ) BHNN; Q % 3AE$H M N I N;E$ ' *
"&'":! )"' '! ' "1+ < " D "
( +,.: Q / Q ) # +
;E$%;%K%A )B $;NH;BA 0H GHAGB 3; H H$% %d$%; _ ;)GH
1* . 9 1" & ' :!+! $%& < )".!" $/ /
- B V < ;)GH -::
;E$%;%K%A )B $;NH;BA 0H GHAGB 3; H H$% %d$%; _ ;)GH
2 '! "&/ ) '& )"., 2 "/!" 2 &P"/ $%& 0
G +
N Q(BH W% W H ? # $ .
A ) 5 K " XH Z. H ) !:
& A!1!#E 4 - .-- 44.--: -:F+
QKHNNHB.0KQ)A;$ 0 M HNNHE)HBG W "/) 1' /! -&') &3
A!+! "&1 &.&+N E = c > V @ ? [ $ -:4+ ! !
A0KQ H ( &.&+" B V < H G C
-:,,
2,-(;HNAK H &.&+" . '":! )" N E = c >< V @ ? $
-:4! -F!
(HBH;B ; Q !: 1 /!1 & 3.& 4) " & '& !) & <5 !& ! 1(.")!
'"3! !1 !) 14:!") '! 1 &2")/& 4 0
_ K 3 ( # .()
(HBH;B ; Q M E0B 0H N M A$% V B Q M 0; $ V Q
B
# & ) -F
2 +-2.+ F
-LKH;BA` V M %BAIeA 0 Q ) Q M AN;IH;B ) M AN;IH;B H $
" $ Q ) 5
( # !:") '! "&.&+" ! "61 " ) ' ! F - !-.
!: F
BA0 N Q V E " &))& "&.&+" ' :!+! $%& <*) ":& <5 ! !/
L* & ( ! ) '! "1+ !/ ! 1 /<* & -:: -:, '% *
K H
$ Q( ;A H I $ ) 3 ; < $ Q( ;A H I $ ) M Q cA $ V M
) B)A$ Q B I 'H * !)L*") <& Q1" 1& '!) "1 ,
B < $ ) 6 ) b$ B
( -::F 2. 2
$ Q( ;A H I $ ) M B UVA H N M $ N H0A ; W M %;H$$HE W
B 5 $
% (H !)L*") + &2! *( " )".!" 22 ! F -.
F2 -:,,
$ E% E V $ ) * * 3" &))& "&.5+" E 2 &'*$%& '!
)! 2".-!" ! " . +!/ '! 1* "!1 !) !/ */ ( ! '! "1+ 1&
! "'5 '& "& 1'! '& & ! ! -,+ '% * K
3 ( # ' *
$ E% E V $ M $AK%A V $ 0
H H $ .BE !:") '! "&.&+" !
"61 " ) ' ! E F 22. + F
$;NI V " &))& "&.&+" ! !. $S!) .&/9 " ) !/ "1+ 1&)
!) '&) ' 4< ! "& 1'! '& & ! ! ,- 3 '% *
K 3 I I
$AK` N M NH;%H W G !+*. $%& ' 2 &'*$%& !/ 3.& !) )
"1!L*"Q1! ) I K3I -::2 -+4
$K0HQ * ."# $%& '& '" +15) " & 3.& !) . '& ) '& ' 4<
!
" #
$ % % & ' "
( ) ( " %* * (
! # + , -. /
-01 ,.23.
4 .!3.
5
' 6
7 4
5 898 :
;
! 6 <
5
5
# 898
" =
=
>
= ?
!
! > = = =
$ % &
% = ' ( = ) ( " %*
* " 4 ! > ≥
- 01 ,.23. >
?
> = 7
) @.!3. A
> =
( 898 >
:
> > =
(
= >
898
4 & ; , CC. ...
B 3 5 D3E 6 !
1.. @4*F)G* $ 3..-H
' $&4&I ) 3..-H 8*J 3..0H :8&8)(K$8< I4 8&( J$4LM<
&4 8<&4N 3..CA
< & #
O "
P Q J
,,E # *
) 4 % % $ J &
% : J &
& 1.. ... R 3
; #
; 5
* @B8NN 3..CA
< % CD 00.
R 3 -S S E 5
S0 E @)TI :4 3..0A
< U O
O !
5 * ;
6
O @' $$ 8$4 ,V11A
;
" @N 4N 3..CA
O
# & @N4:%$ ( ,VV.A
I W * @3..CA 6
O
O 7
#
#
) O
"
< 5
O #
;
# # < #
5 O
@N<% ) 3..-A
@)4&(<)
,VV3A
) @,V1,A O
;
!9 % ' O
;
<
7 %
@,VV3A )
%
5 5
! 5
" ;
O +
5
5
O
< O
!" #$ %&' " (' (#$ %&' ) * " )" "+!,)'
<
' " ( : # 4 N
" ) ( " %
,1 R % I * @,VS3A
! * ) % @)
A ; 3C. <
& ) % BX
; * ! ; O ;
D.. @)TI :4 3..0A 4 !
6 ! ;
- N "
) ( " %
Y " #
I )
! " O
5
< # ;
% ;! <
;
% NT98))<N<) 4$J8))<N<)
& <))<N<) N8(ZN8 <) NT98))<N<) & <))<N<)
N8(ZN8 <) @)TI :4 3..0A
4 4 U @3...A
P # Q [ 8*J
; ; @9 N<)< ,VV,H
8*J ,VV3A
".#/#%&' ) + ( ++"+ )" (!#!,)" ')"( 0"1 /,12 # )" !" "/'
@,VS1A @:I(A
U 5
Y
@:I A ; " I ' @3...A
;
6 7 , 5 H 3
- "
O
4 !
O
< "
01
5 !
4
6 T ( : @T(:A
" ! )4I!DV "
; 8 Y I + @8YIA <
O " )T$' $®@N4&I8: 3..3A
6 7
3 1'+! 0"1 " '("! )" ) )'+
% " # "
" ! ;
O
3. ,. O
@$<I4N ,VV3A
4 5
↔ 4 +
C.. +
,C.
< " !
U " J%) @J % )= A
4 5 5
5 5
'
; 5
; 5 ,
6 ,.
%
6 6
4 # O
5
& O 6 ;
&
C. % 6
+
4 6 ;
@,V-1A %
+ #
& @,VV3A $ @,VV3A '
+ 7 I ' 5\6 I +
] 9 8 + 4
" !
: & 9 3 .0
4 1'+! 0"1 " '("! )" +'('+
% ! 5
+
I ;
,.
%
" .!3.
; :*$4%4 @,VVSA 4 "
! ; @ ;
A @ ;
A @ ;
; A % " 5
^ 3_ : 3_ @ B N!,
A & _ B_ @ : !,
A 4-_ @
B N!, A ^ _ 4 @
. C N!,
A + @
U A
5 !" #$ %&' (#1*!#
I 5
"
; ;
4
3
3 '/)#%6"+ (#1*!# +
< O ;
@3..0!3..CA 5
$%%&!' " ( ; 4
1D- 1 3..0 5
6
@' 3A [ 3..C
, .D- , !
; 6 @' 3A
4 -D V 3..0
-D D 3..C 6
4 3. V
3- 1 3..0 3. 1 30 C 3..C
4 ; S3
E 3..0 01 E
5 [ 3..C
;
6 S. E 0S E 6
6 5
- 9 $%%&!
' " ( ) ( " @%*A
3..0 3..C
3 ( ++"+ )" (!#!,)" / * " "78" #1"/! (
4 ;
4 @' -A
6 5
7 H
" 4 @' 0A
; " 0 50 100 150 200 250 300 350 400 450 500
jan fev mar abr mai jun jul ago set out nov dez
Meses P re ci p it aç ào ( m m ) 0 10 20 30 40 50 60 70 80 U m id ad e re la tiv a d o a r ( % )
Precip 2004 Precip 2005
3- I $%%&!' "
( ) ( " @%*A
4- $ $%%&!
6
7
3CV . 3SS . H 3SS . 31S .
31S . -.C . I
! U
" <
; ,
- I @
; A @
A $%%&!' " (
91:
;/ *7 2)
8&' $8<$ ,, ,3 3, 33 31 3V -. -,
-D -S -1 0- 00 0D 0S 01 3D, . 3SD D 3S. 1
8&( $: I8G$8< , 3 - ,. ,- ,S ,1 ,V 3.
3- 3S -3 -C -V 03 0C 3SS . 31D 1 31, 0
)T% $8<$ 0 C D S 1 V ,0 ,C ,D 30
3C 3D -- -0 0. 0, 31S S -.- C 3V0 0
3 3 /!" 8 "! %&' )'+ ! #<,!'+ .;+# '+ )' +'('
@( 3 - 0A ! 5
5 C13
1S- R !, CD- 133 R !,
D0D 1C, R !, [ O
3C ,0S R !, 11 -,3 R !, H
3S 3.V R !, ,3C -.S R !,
3- ,-D R !, SC 3SV R !,
% &`
J
4 ) 4
!!!!!!!!! R !,!!!!!!!!
$
) a4
I
!-I
!-%
-
!-4
R !,
J '
R !, (
,, S,3 1C 3.- . 03 , 0D 3 1V . 0V ,3S -0S ' !
,3 SDC 0D ,1V . 30 , 0V 3 1- . 0S SC D.- ' !
3, 10V 3C ,3D . 3. , C0 3 V, . 0S C. D.- 4
33 D,D ,,, 3S- . 0, , 3S 3 S1 . C0 ,13 --- ' ! !
31 D1D ,,, 3.- . CC , 0- 3 1. . 01 ,., C.3 ' ! !
3V SCV 1V ,C3 . CV , C, 3 1C . 0S C. DS, ' !
-. S.V V. 3., . 0C , -S 3 SV . C. ,.3 0VC ' ! !
-, S3D VS ,SS . CC , 01 3 V. . 01 ,.. 0-C ' !
-D DDV ,3V 3.3 . D0 , -- 3 S0 . C, ,., C.. ' ! !
-S C13 ,.D -,3 . -0 , 3- 3 DD . C- ,-. CV- ' ! !
-1 S,. 1D 3.0 . 03 , -D 3 1- . C, ,.3 C.. ' ! !
0- D1C ,,3 3.- . CC , -V 3 1, . C. ,,0 0-1 ' ! !
00 1S- -V 11 . 00 , C0 3 V- . 0S 3C S,C 4
0D 133 V. 11 , .3 , 0C 3 1- . 01 3C S,C 4 !
0S D3- ,0S 3-. . D0 , 33 3 S, . C0 ,SV 33, ' ! !
01 SCV 1V ,C3 . CV , 0. 3 V. . C, SD C.. ' !
3- 4 $%%&!' " (
% &`
J
4 ) 4
!!!!!!!!! R !,!!!!!!!!
$
) a4
I
!-I
!-%
-
!-4
R !,
J '
R !, (
., CD- 3.V 331 . V3 , 03 3 V0 . C, ,3D 00S ' ! !
.3 S-D ,-1 ,3D , ,. , 00 3 13 . 01 SC 0.0 ' !
.- DS3 ,SS ,C, , ,S , 00 3 S1 . 01 SC C.- ' !
,. S-C ,,0 ,C, . SC , 0V 3 1. . 0D ,.. --S ' !
,- SSV 0C ,SD . 3D , 01 3 10 . 0S SC C-S ' !
,S S.. VV 3., . 0V , 0D 3 1S . 0V ,3D -S- ' ! !
,1 133 3S ,C, . ,1 , CD 3 SV . 00 SC C.- 4
,V S1- V, ,3D . S3 , C- 3 1C . 0D SD -VD 4
3. S01 S0 ,S1 . 03 , 0D 3 1. . 0S ,., 0-3 ' !
3- S0, 13 ,SS . 0D , C3 3 1V . 0S ,., 03V ' !
3S 1.- CV ,-1 . 0- , C- 3 1S . 0D C. D-S 4
-3 SSD S3 ,C3 . 0S , 0D 3 1C . 01 ,., --C 4
-C SV1 SS ,3C . D3 , 01 3 10 . 0S C. D.. 4
-V C10 ,.V -.S . -D , -S 3 S0 . C. ,C- C., 4 !
03 SD0 1C ,C, . CD , C. 3 11 . 0S 11 0,S 4 !
4- 4 $%%&!' " (
% &`
J
4 ) 4
!!!!!!!!! R !,!!!!!!!!
$
) a4
I
!-I
!-%
-
!-4
R !,
J '
R !, (
.0 S.. ,3- ,SS . DV , 0C 3 13 . 01 SC CSD 4 !
.C 1CC S. SC . V- , C- 3 11 . 0D 3C DDD 4 !
.D SSV 13 ,-V . CV , 01 3 1S . 01 C. D0. ' !
.S SD- ,-D ,., , -C , 0V 3 SV . 0D C. C.0 ' !
.1 SS. ,.0 ,3D . 1- , C. 3 1- . 0D SC 0.0 ' !
.V DVD ,C3 ,C3 , .. , 0V 3 1D . 0S ,., --C ' !
,0 S-3 DD 3.3 . -- , 0D 3 1- . 01 ,., 0VC ' ! !
,C S-. VC ,SC . C0 , C0 3 V3 . 0S ,.. 031 ' !
,D 1C, 3- ,3D . ,1 , 0- 3 S0 . 0S C. D.- 4 !
30 S-, V, ,S1 . C, , 0- 3 10 . 0V ,., 0-3 4 !
3C 1-3 D1 ,.. . D1 , CV 3 V3 . 0C C. C.. 4 !
3D SV, S, ,-1 . C, , CS 3 V- . 0D C. D-S ' !
-- D0D SD 3SV . 3S , 0, 3 V0 . C, ,SS -DC ' ! !
-0 SDD ,.V ,3C . 1S , CD 3 V3 . 0D SC 0.. ' !
0. SD. 11 ,C3 . C1 , 0- 3 1C . 0V ,., --C ' !
4 7
! ! !
+ ! ! &
7 !
! ! ! 5 ,D
! % U
!
4 "
" ? R @,VD3A " !
a <
5 5 a ; 5 . ,CH ;
" < ! 5
a
5
"
O 5 ; NT98))<N<
& <))<N<)
< ! , 33
, C0 !- 3 V, 3 V-
!-H
, -S , CD !- 3 S0
3 V0 !- , -D , CV
!- 3 S0 3 V0
!-(
5 ; 3 DC
!-< 6
!
. 00 . C0 -
Y 3C ,13 R !,
C. ,C- R !, 3C ,SS R !,
3 4 /!" 8 "! %&' )'+ ! #<,!'+ =,;1# '+ )' +'('
) : @3..0A 5 !
5 ;
5 # "
< ; 5
C @4& b<) , 3 -A
5- 9 ; 5 c
c 4 N # 4 ) G 4 ;
' ) !%* 3..S
^ ! 5 6
"
< # ;
@' C A 4
# ; "
5 5
" O
<
H ;
8> ?@ @ @ 0@ >@@ (@
^3< - !!!!!!!!!!!!!!!!!!!!!!!!!!! !-!!!!!!!!!!!!!!!!!!!!!!! E R !,
D 0V ,1 . . 0 . , C 1 , D - . S V ,. 1 S, -- 3
D 0D ,. 0 . - . , 0 D , - 3 V D 0 V - D1 ,D V
-@' C A N @,VVSA 5
! +
U
< ! ;
@' C C A ) @3..DA
5- 9 ; % @ A B_ @ A _3 @ A : _3
@ A
4
5 D 0 S V
!-V - ,. 1 !- D1 S3 E @' D D D A
& ! 5 C.E
#
< U ; + ;
0,0 2,0 4,0 6,0 8,0 10,0 12,0 14,0 16,0 18,0 20,0
INFERIOR INTERMEDIÁRIO SUPERIOR
Níveis de altitude
P ( m d m 3 ) 0,0 0,1 0,2 0,3 0,4 0,5
INFERIOR INTERMEDIÁRIO SUPERIOR
Níveis de altitude
K + ( cm o lc d m 3) 0,0 1,0 2,0 3,0 4,0 5,0 6,0 7,0
INFERIOR INTERMEDIÁRIO SUPERIOR
Níveis de altitude
C
a
+
2 (
cm o l c d m 3 ) 0,0 0,2 0,4 0,6 0,8 1,0 1,2 1,4 1,6 1,8
INFERIOR INTERMEDIÁRIO SUPERIOR
Níveis de altitude
M
g
+
3 (
cm o lc d m 3) b a
@' D A J O ! !
; +
5
5 +
A- 9 ; )* @ A ( @ A 9 @ A :< @ A
&
5 #
< ! 5
5
;
4
O
@$ ) &I ,VVCH *<($ N 3..3A
0,0 1,0 2,0 3,0 4,0 5,0 6,0 7,0 8,0
INFERIOR INTERMEDIÁRIO SUPERIOR
Níveis de altitude
S B ( cm o l c d m -3) 8,0 8,5 9,0 9,5 10,0 10,5 11,0
INFERIOR INTERMEDIÁRIO SUPERIOR Níveis de altitude
C T C ( cm o lc d m -3) 0 10 20 30 40 50 60 70 80 90 100
INFERIOR INTERMEDIÁRIO SUPERIOR Níveis de altitude
V ( % ) 0,0 5,0 10,0 15,0 20,0 25,0 30,0 35,0
INFERIOR INTERMEDIÁRIO SUPERIOR Níveis de altitude
M O ( g k g -1 )
a b
3 5 +! ,!, B' #$'/! (
< + #
" 6