Sidnei Eduardo Lima-Junior*, Isabella Braz Cardone and Roberto Goitein
Departamento de Zoologia, Instituto de Biociências, Universidade Estadual Paulista, Av. 24-A, 1515, 13506-900, Rio Claro, São Paulo, Brasil. *Author for correspondence.
! "
## $ % & " ! !! # #
!! ! ## " # $ ! !
% & # # # # ! " # & !
! " ! ! # ! " #
# ! ' ( # ' &
" # & # # !! ! ! ##
# # ! ## " #
!
! ) # & #
!"#! # $
# ! % * " # + ", -. )
/ # / $ % . "
+ # # ! # # 0 !
" $ -1 ) # % +
# # # " ' -. + #
/ + " # ! + # 0 -. #
" -1 # ( ' -. ' #
. + / + # # !
2 # 0 # " /
+ -. )
& # ' (!) ' / ! -. 2 -. # -.
' ( ! + # )
! ! " # !
! # ! # # "
# # ! #
) " & "
3 )
" #
# # ! !
3 !
# # + # !
& # # $3
4 567 3 # ' ## 4 5 %
# # '
( $ % & ## " " )
& &
# # # $8 '
4 947 : ; 4 97 3 4 567 3 #
' ## 4 5 % & <
# #
! # " ! #
& 0 " ! )
& # ! #
& # ! , ! !
# ! &
! ! #
# !
! ## '
( " ! )
& " # &
# + $ % & ,
! ! # # = &
! & " &
! # # !
!! ! #
+ & # ; ## # #
*+, - (.
4 % ? # # #
" # ! #
! $ 3 " 4 %
## $' 4 %
' "#
&
# # !! &
" !
! #! ## # #
& # ! ! !
$ # ! 8 %
! ' " 2 : $ @4 A AA
@ A AAB% ! . C # 3 0 #
" & D * "
8 $ 4 % # B $
4 % & " ! ##
E ! ! #
# 8 ## # "
# ! ) ! #
# & ! $
" # , !
# # '
( % & # !
# ## 44 8 &
! # 0 # ! "
# & "# 4 #
! ! " # ! "#
8 #
B # & "#
/# 0 F " $ % # & ! # 0
# ! # " #
# $ % # & $ %
4 5 4 4 6
& 5 4 4 9
5 4 4 9
45 5 4 4 9 46 9 94
& ! #
# +
# & " $!
) % & & G $4%
## 9 # # $ %
! ##
' ( $ % & # # "
" " & : #
! ) $ "#
8 # B % &
! H ; # B ## &
# #
$' 4 5 % αI 9
( 4 "# #
" # & ! !
8 !! : # B
& ## ## #
& !
, + ! # & #
! ## # ! ## & G
B I 54 8 5
+ ! # & #
, # !
! ## & G
G B I 8
& G B I 9 8 69
G B I 5 5 8 9 6
G B I 8 949
* ( "# &
' ( " ! ! &
# ! ## !
& ? !
# ! # !!
! ! ! " & !
!
quartile range (25-75%) median
S
ta
n
d
a
rd
L
e
n
g
th
(
c
m
)
8.4 8.8 9.2 9.6 10 10.4 10.8
0 10.0
autumn winter spring summer A
quartile range (25-75%) median
T
o
ta
l
W
e
ig
h
t
(g
)
15 22.5 30 37.5 45
0 30.0 45.0
autumn winter spring summer B
$ 1 0 + # ! 8 $ %
) 2 ! #! # 2 ## ! $ ! *++ quartile range (25-75%) median 4.2 4.4 4.6 4.8 5 5.2 0
autumn winter spring summer
(b=2.833 for all seasons)
5.0 A C o n d it io n F a c to r ( K .1 0 2 ) quartile range (25-75%) median 2 3.6 5.2 6.8 8.4 10 0 B
autumn winter spring summer
(b=3.040) (b=2.653) (b=2.596) (b=2.515)
10.0 C o n d it io n F a c to r ( K .1 0 2 )
$ 1 3 + # ! #
" # ! ' ( E ) &
# # + # ! ! ! # "
# E 3 # ! " #
/# 3 # $H ; # B ## % !
8 # B ' ( $&
# % " # $αI 9%
8 # B
' ( $ % ' ( $ # %
& 9 $ % 9 $ % 9 $ % 9 $J%
9 $ % 9 $ % 9 $ % 9 $J%
9 $ % 9 $J% 9 $J% 9 $J%
B 9 $ % 9 $ % 9 $ % 9 $J%
B 9 $ % 9 $J% 9 $J% 9 $J%
9 $ % 9 $J% 9 $J% 9 $ %
G ! !! 7 J ! !!
# ! ) # #
## ' ( $ %
$ % & $ %
# " # # ' + #
& ! # &
# ' ( $3 #
' # 4 5 % (
# # # &
## # ' (
# ! ## & & !
#
: # & ! "
# & !! &
# ! ! +
) # #
& & !
& # &
"
! " ## ) # !
!
? " # # &
# & & & # # &
! #
' + # ! #
& $ % ! #
## ##
# & # B !! $ % & #
! ' (
) ! #
+ ! # &
# ) "
# & #
' " #
# "#
)
! " #
! ! !! # !
" # # ) ) #
# ! ## ! #
# & ## "
# # !! ! #
& # # !
# ! " #
! # &
# $ % ## + #
' ( $3 4 56 4 %
& " #
& # , ! "
# # ! "#
# ! " !
& " + ! # &
# & , # "
## + # !!
$ % ' ( &
( 3
E & " #
# # " " !
# & # " " ##
# #
# # !! ! # E
& & )
" 0 !
#& ! ! $ %
"
# # ! # ! "
# # " # "
= & # "
0 !
455 - (.
" # !! " # #
# ! " # &
# & #
! # !
! !
' (
!! # #
! + "# # ; ##
# & # ! ## " #
$ # ##
" # ! ) # %
" ( CK C 'FC+
F*:E F E ( 7 3 :3EK:E L : # -. # M
# ! -.
8 4 66 $ # !
% . N E C:
4 # 4 4 4 4
F ?FEO E * ! # P
# ! -.
: 4 5 $? 8 %
D . C # 3 # ! " # "
. C # 4 5 4 4 59
3 :3EK:E L 7 >K: FE D : ? ! -.
2 !!
$8 4 95% $? 8 %
, # $ . ' # C% $ $
. C # 699 695 4 5
3?8LK: 7 '?FF?88Q C 8 # !
"# ! # ! !
% & ! 4 4
45 4 5
3: L ( K ! -. # -.
M # ) '
! ! : D 6 6 4 56
3: L ( # ! -.
$C ! % '
49 449 4
' C? = ( ) * + ,
C " G K 8R 4 5
' : D D ' * S -.
- . " K 4 5 $C
' % : E" K . C #
3 # ! " # " . C # 54
59 4
8K ':KF K * # & #
# # &
$ / % % 0 ( ?)! 4 4
4 94
:E'HK: B K ' !
" # # ! ! # ! & ' !"
$ ? & 4 4 4 5 4 9
K:LE?O H E " #
! # ! 0 # ;
! " ! $ % & !
6 9 66 4
